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CONDENSED CONSOLIDATED STATEMENTS OF CASH FLOWS - USD ($)
$ in Thousands
 3 Months Ended
Mar. 31, 2025
Mar. 31, 2024
Operating Activities:    
Net loss $ (30,597) $ (49,089)
Adjustments to reconcile net loss to net cash used in operating activities:    
Depreciation and amortization 1,020 1,406
Amortization in operating lease right-of-use assets 904 1,120
Stock-based compensation 2,596 2,719
Impairment of long-lived assets 0 4,349
Accretion of discount on marketable securities 0 (273)
Deferred income tax benefit 15 0
Other (10) 0
Net changes in operating assets and liabilities:    
Accounts receivable 160 232
Prepaid expenses and other assets (2,631) 962
Refundable research income tax credits 0 (1,391)
Accounts payable and other accrued liabilities 4,216 (4,939)
Accrued compensation and employee benefits (591) (2,559)
Lease liabilities (1,231) (1,218)
Other non-current liabilities 0 18
Net cash used in operating activities (26,149) (48,663)
Investing Activities:    
Maturities of marketable securities 0 1,110
Proceeds from sale of marketable securities 0 34,730
Net cash provided by investing activities 0 35,840
Financing Activities:    
Proceeds from at-the-market offering, net of offering expenses 10,361 0
Proceeds from issuance of common stock, net of offering expenses 0 22,524
Taxes paid related to net share settlement of equity awards (2,227) (402)
Net cash provided by financing activities 8,134 22,122
Effect of exchange rate changes on cash, cash equivalents, and restricted cash 1,277 (670)
Net (decrease) increase in cash, cash equivalents, and restricted cash (16,738) 8,629
Cash, cash equivalents, and restricted cash, beginning of period 43,418 46,704
Cash, cash equivalents, and restricted cash, end of period 26,680 55,333
Supplemental cash flow disclosures:    
Property and equipment included in unpaid liabilities 234 460
Offering expenses in relation to issuance of common stock in at-the-market offering included in unpaid liabilities 165 0
Offering expenses in relation to issuance of common stock included in unpaid liabilities $ 0 $ 736