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CONDENSED CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY - USD ($) $ in Thousands | Total | At-The-Market Offering | Stock Issuances Excluding At-The-Market Offering | Common Stock | Common Stock At-The-Market Offering | Common Stock Stock Issuances Excluding At-The-Market Offering | Additional Paid-in Capital | Additional Paid-in Capital At-The-Market Offering | Additional Paid-in Capital Stock Issuances Excluding At-The-Market Offering | Accumulated Deficit | Accumulated Other Comprehensive Loss
--- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | ---
Beginning balance (in shares) at Dec. 31, 2023 |  |  |  | 178,133,548 |  |  |  |  |  |  |
Beginning balance at Dec. 31, 2023 | $ 82,887 |  |  | $ 1,781 |  |  | $ 1,492,077 |  |  | $ (1,406,376) | $ (4,595)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock (in shares) |  |  |  |  |  | 24,761,905 |  |  |  |  |
Issuance of common stock |  |  | $ 21,788 |  |  | $ 248 |  |  | $ 21,540 |  |
Issuance of common stock upon exercise of pre-funded warrants (in shares) |  |  |  | 3,809,523 |  |  |  |  |  |  |
Issuance of common stock upon exercise of pre-funded warrants | 71 |  |  | $ 38 |  |  | 33 |  |  |  |
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax (in shares) |  |  |  | 1,116,678 |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax | (481) |  |  | $ 11 |  |  | (492) |  |  |  |
Issuance of common stock under employee stock purchase plan (in shares) |  |  |  | 379,486 |  |  |  |  |  |  |
Issuance of common stock under employee stock purchase plan | 146 |  |  | $ 4 |  |  | 142 |  |  |  |
Stock-based compensation | 5,784 |  |  |  |  |  | 5,784 |  |  |  |
Foreign currency translation adjustment | (1,293) |  |  |  |  |  |  |  |  |  | (1,293)
Net pension gains | 238 |  |  |  |  |  |  |  |  |  | 238
Unrealized loss on marketable securities, net of tax | (233) |  |  |  |  |  |  |  |  |  | (233)
Net loss | (85,217) |  |  |  |  |  |  |  |  | (85,217) |
Ending balance (in shares) at Jun. 30, 2024 |  |  |  | 208,201,140 |  |  |  |  |  |  |
Ending balance at Jun. 30, 2024 | 23,690 |  |  | $ 2,082 |  |  | 1,519,084 |  |  | (1,491,593) | (5,883)
Beginning balance (in shares) at Mar. 31, 2024 |  |  |  | 203,686,086 |  |  |  |  |  |  |
Beginning balance at Mar. 31, 2024 | 56,697 |  |  | $ 2,037 |  |  | 1,515,926 |  |  | (1,455,465) | (5,801)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock upon exercise of pre-funded warrants (in shares) |  |  |  | 3,809,523 |  |  |  |  |  |  |
Issuance of common stock upon exercise of pre-funded warrants | 71 |  |  | $ 38 |  |  | 33 |  |  |  |
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax (in shares) |  |  |  | 326,045 |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax | (79) |  |  | $ 3 |  |  | (82) |  |  |  |
Issuance of common stock under employee stock purchase plan (in shares) |  |  |  | 379,486 |  |  |  |  |  |  |
Issuance of common stock under employee stock purchase plan | 146 |  |  | $ 4 |  |  | 142 |  |  |  |
Stock-based compensation | 3,065 |  |  |  |  |  | 3,065 |  |  |  |
Foreign currency translation adjustment | (313) |  |  |  |  |  |  |  |  |  | (313)
Net pension gains | 231 |  |  |  |  |  |  |  |  |  | 231
Unrealized loss on marketable securities, net of tax | 0 |  |  |  |  |  |  |  |  |  |
Net loss | (36,128) |  |  |  |  |  |  |  |  | (36,128) |
Ending balance (in shares) at Jun. 30, 2024 |  |  |  | 208,201,140 |  |  |  |  |  |  |
Ending balance at Jun. 30, 2024 | 23,690 |  |  | $ 2,082 |  |  | 1,519,084 |  |  | (1,491,593) | (5,883)
Beginning balance (in shares) at Dec. 31, 2024 |  |  |  | 212,837,679 |  |  |  |  |  |  |
Beginning balance at Dec. 31, 2024 | 22,770 |  |  | $ 2,128 |  |  | 1,532,489 |  |  | (1,504,317) | (7,530)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock (in shares) |  |  |  |  | 20,807,709 | 12,235,000 |  |  |  |  |
Issuance of common stock |  | $ 17,627 | 21,139 |  | $ 209 | $ 122 |  | $ 17,418 | 21,017 |  |
Issuance of common stock upon exercise of pre-funded warrants (in shares) |  |  |  | 22,398,393 |  |  |  |  |  |  |
Issuance of common stock upon exercise of pre-funded warrants | 224 |  |  | $ 224 |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax (in shares) |  |  |  | 4,061,971 |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax | (2,647) |  |  | $ 41 |  |  | (2,688) |  |  |  |
Issuance of common stock under employee stock purchase plan (in shares) |  |  |  | 463,135 |  |  |  |  |  |  |
Issuance of common stock under employee stock purchase plan | 179 |  |  | $ 4 |  |  | 175 |  |  |  |
Stock-based compensation | 4,822 |  |  |  |  |  | 4,822 |  |  |  |
Foreign currency translation adjustment | 5,980 |  |  |  |  |  |  |  |  |  | 5,980
Net pension gains | 91 |  |  |  |  |  |  |  |  |  | 91
Unrealized loss on marketable securities, net of tax | 0 |  |  |  |  |  |  |  |  |  |
Net loss | (50,583) |  |  |  |  |  |  |  |  | (50,583) |
Ending balance (in shares) at Jun. 30, 2025 |  |  |  | 272,803,887 |  |  |  |  |  |  |
Ending balance at Jun. 30, 2025 | 19,602 |  |  | $ 2,728 |  |  | 1,573,233 |  |  | (1,554,900) | (1,459)
Beginning balance (in shares) at Mar. 31, 2025 |  |  |  | 225,364,090 |  |  |  |  |  |  |
Beginning balance at Mar. 31, 2025 | 4,906 |  |  | $ 2,254 |  |  | 1,543,093 |  |  | (1,534,914) | (5,527)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock (in shares) |  |  |  |  | 11,278,957 | 12,235,000 |  |  |  |  |
Issuance of common stock |  | $ 7,266 | $ 21,139 |  | $ 113 | $ 122 |  | $ 7,153 | $ 21,017 |  |
Issuance of common stock upon exercise of pre-funded warrants (in shares) |  |  |  | 22,398,393 |  |  |  |  |  |  |
Issuance of common stock upon exercise of pre-funded warrants | 224 |  |  | $ 224 |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax (in shares) |  |  |  | 1,064,312 |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax | (420) |  |  | $ 11 |  |  | (431) |  |  |  |
Issuance of common stock under employee stock purchase plan (in shares) |  |  |  | 463,135 |  |  |  |  |  |  |
Issuance of common stock under employee stock purchase plan | 179 |  |  | $ 4 |  |  | 175 |  |  |  |
Stock-based compensation | 2,226 |  |  |  |  |  | 2,226 |  |  |  |
Foreign currency translation adjustment | 4,068 |  |  |  |  |  |  |  |  |  | 4,068
Net pension gains | 0 |  |  |  |  |  |  |  |  |  |
Unrealized loss on marketable securities, net of tax | 0 |  |  |  |  |  |  |  |  |  |
Net loss | (19,986) |  |  |  |  |  |  |  |  | (19,986) |
Ending balance (in shares) at Jun. 30, 2025 |  |  |  | 272,803,887 |  |  |  |  |  |  |
Ending balance at Jun. 30, 2025 | $ 19,602 |  |  | $ 2,728 |  |  | $ 1,573,233 |  |  | $ (1,554,900) | $ (1,459)

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