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CONDENSED CONSOLIDATED STATEMENTS OF CASH FLOWS - USD ($)
$ in Thousands
 6 Months Ended
Jun. 30, 2025
Jun. 30, 2024
Operating Activities:    
Net loss $ (50,583) $ (85,217)
Adjustments to reconcile net loss to net cash used in operating activities:    
Depreciation and amortization 2,013 2,648
Amortization in operating lease right-of-use assets 1,808 2,451
Stock-based compensation 4,822 5,784
Impairment of long-lived assets 0 5,521
Accretion of discount on marketable securities 0 (273)
Other 35 (103)
Net changes in operating assets and liabilities:    
Interest receivable 0 370
Accounts receivable 154 342
Prepaid expenses and other assets (1,389) 2,433
Refundable research income tax credits 0 1,132
Accounts payable and other accrued liabilities 301 (4,654)
Accrued compensation and employee benefits 764 (3,195)
Lease liabilities (2,262) (2,822)
Other non-current liabilities 30 36
Net cash used in operating activities (44,307) (75,547)
Investing Activities:    
Maturities of marketable securities 0 1,110
Proceeds from sale of marketable securities 0 34,730
Proceeds from sale of assets classified as held for sale 0 127
Purchases of property and equipment (24) 0
Net cash (used in) provided by investing activities (24) 35,967
Financing Activities:    
Proceeds from issuance of common stock, net of offering expenses 21,589 21,924
Proceeds from at-the-market offering, net of offering expenses 17,627 0
Taxes paid related to net share settlement of equity awards (2,647) (481)
Proceeds from issuance of common stock under employee stock purchase plan 179 146
Net cash provided by financing activities 36,748 21,589
Effect of exchange rate changes on cash, cash equivalents, and restricted cash 4,009 (927)
Net decrease in cash, cash equivalents, and restricted cash (3,574) (18,918)
Cash, cash equivalents, and restricted cash, beginning of period 43,418 46,704
Cash, cash equivalents, and restricted cash, end of period 39,844 27,786
Supplemental cash flow disclosures:    
Property and equipment included in unpaid liabilities 101 433
Offering expenses in relation to issuance of common stock included in unpaid liabilities $ 226 $ 0