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Condensed Consolidated Statements of Stockholders' Equity (Unaudited) - USD ($) $ in Thousands | Total | Pfizer License Agreement | Common Stock | Common Stock Pfizer License Agreement | Additional Paid-in Capital | Additional Paid-in Capital Pfizer License Agreement | Accumulated Other Comprehensive (Loss) Income | Accumulated Deficit
--- | --- | --- | --- | --- | --- | --- | --- | ---
Beginning balance at Dec. 31, 2022 | $ 160,824 |  | $ 34 |  | $ 373,225 |  |  | $ (212,435)
Beginning Balance, Shares at Dec. 31, 2022 |  |  | 34,958,730 |  |  |  |  |
Issuance of common stock |  | $ 5,000 |  | $ 2 |  | $ 4,998 |  |
Issuance of common stock (Shares) |  |  |  | 1,811,594 |  |  |  |
Issuance of restricted common stock, net of tax withholdings | 2 |  | $ 1 |  | 1 |  |  |
Issuance of restricted common stock, net of tax withholdings, (Shares) |  |  | 148,047 |  |  |  |  |
Stock-based compensation | 4,884 |  |  |  | 4,884 |  |  |
Net unrealized (loss) gain on marketable debt securities | 696 |  |  |  |  |  | $ 696 |
Net loss | (19,243) |  |  |  |  |  |  | (19,243)
Ending balance at Mar. 31, 2023 | 152,163 |  | $ 37 |  | 383,108 |  | 696 | (231,678)
Ending balance (in shares) at Mar. 31, 2023 |  |  | 36,918,371 |  |  |  |  |
Beginning balance at Dec. 31, 2022 | 160,824 |  | $ 34 |  | 373,225 |  |  | (212,435)
Beginning Balance, Shares at Dec. 31, 2022 |  |  | 34,958,730 |  |  |  |  |
Shares issued pursuant to the ATM program, net of commission (shares) |  |  | 1,001,208 |  |  |  |  |
Net unrealized (loss) gain on marketable debt securities | (85) |  |  |  |  |  |  |
Net loss | (35,144) |  |  |  |  |  |  |
Ending balance at Jun. 30, 2023 | 145,275 |  | $ 39 |  | 392,900 |  | (85) | (247,579)
Ending balance (in shares) at Jun. 30, 2023 |  |  | 39,376,941 |  |  |  |  |
Beginning balance at Mar. 31, 2023 | 152,163 |  | $ 37 |  | 383,108 |  | 696 | (231,678)
Beginning Balance, Shares at Mar. 31, 2023 |  |  | 36,918,371 |  |  |  |  |
Vesting of restricted common stock (Shares) |  |  | 1,436,866 |  |  |  |  |
Vesting of restricted common stock | 1 |  | $ 1 |  |  |  |  |
Shares issued pursuant to the ATM program, net of commission (shares) |  |  | 1,001,208 |  |  |  |  |
Shares issued pursuant to the ATM program, net of commission | 6,122 |  | $ 1 |  | 6,121 |  |  |
Stock options exercised | 3 |  |  |  | 3 |  |  |
Stock options exercised (Shares) |  |  | 20,496 |  |  |  |  |
Stock-based compensation | 3,668 |  |  |  | 3,668 |  |  |
Net unrealized (loss) gain on marketable debt securities | (781) |  |  |  |  |  | (781) |
Net loss | (15,901) |  |  |  |  |  |  | (15,901)
Ending balance at Jun. 30, 2023 | 145,275 |  | $ 39 |  | 392,900 |  | (85) | (247,579)
Ending balance (in shares) at Jun. 30, 2023 |  |  | 39,376,941 |  |  |  |  |
Beginning balance at Dec. 31, 2023 | 125,704 |  | $ 45 |  | 411,821 |  | 63 | (286,225)
Beginning Balance, Shares at Dec. 31, 2023 |  |  | 44,754,853 |  |  |  |  |
Issuance of common stock in private placement, net of offering costs (See Note 7) (Shares) |  |  | 8,849,371 |  |  |  |  |
Issuance of common stock in private placement, net of offering costs (See Note 7) | 39,172 |  | $ 9 |  | 39,163 |  |  |
Issuance of common stock pursuant to at-the-market program, net of offering costs (See Note 7) (Shares) |  |  | 3,600,000 |  |  |  |  |
Issuance of common stock pursuant to at-the-market program, net of offering costs (See Note 7) | 10,590 |  | $ 4 |  | 10,586 |  |  |
Issuance of pre-funded warrants in private placement, net of offering costs (See Note 8) | 7,700 |  |  |  | 7,700 |  |  |
Issuance of restricted common stock, net of tax withholdings | (196) |  | $ 1 |  | (197) |  |  |
Issuance of restricted common stock, net of tax withholdings, (Shares) |  |  | 1,497,921 |  |  |  |  |
Stock options exercised | 245 |  |  |  | 245 |  |  |
Stock options exercised (Shares) |  |  | 100,981 |  |  |  |  |
Stock-based compensation | 4,320 |  |  |  | 4,320 |  |  |
Net unrealized (loss) gain on marketable debt securities | (123) |  |  |  |  |  | (123) |
Net loss | (3,256) |  |  |  |  |  |  | (3,256)
Ending balance at Mar. 31, 2024 | 184,156 |  | $ 59 |  | 473,638 |  | (60) | (289,481)
Ending balance (in shares) at Mar. 31, 2024 |  |  | 58,803,126 |  |  |  |  |
Beginning balance at Dec. 31, 2023 | $ 125,704 |  | $ 45 |  | 411,821 |  | 63 | (286,225)
Beginning Balance, Shares at Dec. 31, 2023 |  |  | 44,754,853 |  |  |  |  |
Shares issued pursuant to the ATM program, net of commission (shares) |  |  | 3,600,000 |  |  |  |  |
Stock options exercised (Shares) | 122,551 |  |  |  |  |  |  |
Net unrealized (loss) gain on marketable debt securities | $ (165) |  |  |  |  |  |  |
Net loss | (20,557) |  |  |  |  |  |  |
Ending balance at Jun. 30, 2024 | 169,794 |  | $ 59 |  | 476,619 |  | (102) | (306,782)
Ending balance (in shares) at Jun. 30, 2024 |  |  | 58,914,712 |  |  |  |  |
Beginning balance at Mar. 31, 2024 | 184,156 |  | $ 59 |  | 473,638 |  | (60) | (289,481)
Beginning Balance, Shares at Mar. 31, 2024 |  |  | 58,803,126 |  |  |  |  |
Issuance of common stock under employee stock purchase plan ("ESPP") | 105 |  |  |  | 105 |  |  |
Issuance of common stock under employee stock purchase plan ("ESPP") (Shares) |  |  | 59,882 |  |  |  |  |
Issuance of restricted common stock, net of tax withholdings | (43) |  |  |  | (43) |  |  |
Issuance of restricted common stock, net of tax withholdings, (Shares) |  |  | 30,553 |  |  |  |  |
Stock options exercised | 48 |  |  |  | 48 |  |  |
Stock options exercised (Shares) |  |  | 21,151 |  |  |  |  |
Stock-based compensation | 2,871 |  |  |  | 2,871 |  |  |
Net unrealized (loss) gain on marketable debt securities | (42) |  |  |  |  |  | (42) |
Net loss | (17,301) |  |  |  |  |  |  | (17,301)
Ending balance at Jun. 30, 2024 | $ 169,794 |  | $ 59 |  | $ 476,619 |  | $ (102) | $ (306,782)
Ending balance (in shares) at Jun. 30, 2024 |  |  | 58,914,712 |  |  |  |  |

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