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Income Taxes - Significant Components of Deferred Tax Assets and Liabilities (Details) - USD ($) $ in Thousands | Dec. 31, 2017 | Dec. 31, 2016
--- | --- | ---
Deferred tax assets: |  |
Capitalized research expense | $ 8,546 | $ 567
Net operating loss carryforwards | 71,665 | 95,500
Research and development and other tax credits | 7,531 | 5,300
Deferred revenue | 297 | 5,452
Deferred rent | 2,097 | 2,231
Stock-based compensation | 3,091 | 4,511
Acquired intangibles | 858 | 989
Other | 1,906 | 3,783
Deferred tax assets, gross | 95,991 | 118,333
Valuation allowance | (94,039) | (113,407)
Total deferred tax assets | 1,952 | 4,926
Deferred tax liabilities: |  |
Acquired intangibles | (124) | (175)
Investment in affiliated entity | (422) | (3,624)
Fixed assets | (1,430) | (1,302)
Net deferred tax liabilities | $ (24) | $ (175)

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