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Impact of Recently Issued Accounting Standards - Impact of Adopting ASU 2014-09 (Details) - USD ($) | 3 Months Ended
--- | ---
Mar. 31, 2018 | Mar. 31, 2017
Revenues [Abstract] |  |
Revenue under collaborative research and development arrangements | $ 1,289,046 | $ 4,288,586
Revenue under collaborative research and development arrangements with affiliated entities | 148,008 | 233,330
Grants and miscellaneous revenue | 92,590 | 5,240,233
Grants and miscellaneous revenue from affiliated entity | 0 | 614,036
Total revenues | 1,529,644 | 10,376,185
Operating Expenses [Abstract] |  |
Research and development | 24,577,751 | 24,542,504
General and administrative | 9,698,015 | 7,767,589
Total operating expenses | 34,275,766 | $ 32,310,093
Calculated under Revenue Guidance in Effect before Topic 606 |  |
Revenues [Abstract] |  |
Revenue under collaborative research and development arrangements | 1,289,046 |
Revenue under collaborative research and development arrangements with affiliated entities | 148,008 |
Grants and miscellaneous revenue | 1,850,341 |
Grants and miscellaneous revenue from affiliated entity | 464,400 |
Total revenues | 3,751,795 |
Operating Expenses [Abstract] |  |
Research and development | 22,355,600 |
General and administrative | 9,698,015 |
Total operating expenses | 32,053,615 |
Accounting Standards Update 2014-09 | Difference between Revenue Guidance in Effect before and after Topic 606 |  |
Revenues [Abstract] |  |
Revenue under collaborative research and development arrangements | 0 |
Revenue under collaborative research and development arrangements with affiliated entities | 0 |
Grants and miscellaneous revenue | (1,757,751) |
Grants and miscellaneous revenue from affiliated entity | (464,400) |
Total revenues | (2,222,151) |
Operating Expenses [Abstract] |  |
Research and development | (2,222,151) |
General and administrative | 0 |
Total operating expenses | $ (2,222,151) |

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