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Collaborative Agreements (Details) - Collaborative Arrangement, Product |  |  |  | 1 Months Ended | 3 Months Ended
--- | --- | --- | --- | --- | ---
Dec. 31, 2017 USD ($) | Dec. 29, 2017 | Aug. 07, 2015 product | Mar. 31, 2018 USD ($) | Dec. 31, 2017 USD ($) | Feb. 28, 2017 USD ($) | Sep. 30, 2015 USD ($) | Apr. 30, 2015 USD ($) | Sep. 30, 2013 USD ($) | Mar. 31, 2018 USD ($) | Mar. 31, 2017 USD ($)
ApolloBio |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |  |  |  |  |
Territory expansion option period |  | 3 years |  |  |  |  |  |  |  |  |
Proceeds received in upfront payment |  |  |  | $ 19,400,000 |  |  |  |  |  |  |
Upfront payment received |  |  |  | 23,000,000 |  |  |  |  |  | $ 23,000,000 |
Advisory fees |  |  |  | 960,000 |  |  |  |  |  | 960,000 |
Additional revenue to be achieved |  |  |  |  |  |  |  |  |  | $ 20,000,000 |
Royalty period |  |  |  |  |  |  |  |  |  | 10 years |
Period from effective date for termination |  |  |  |  |  |  |  |  |  | 1 year |
Number of days written notice defore termination |  |  |  |  |  |  |  |  |  | 90 days |
MedImmune |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |  |  |  |  |
Upfront payment received | $ 27,500,000 |  |  |  |  |  | $ 27,500,000 |  |  |  |
Anticipated development and regulatory event based payment receivable milestones |  |  |  |  |  |  |  |  |  | $ 355,000,000 |
Anticipated commercial event based payment receivable milestones |  |  |  |  |  |  |  |  |  | 345,000,000 |
Number of additional products to be developed | product |  |  | 2 |  |  |  |  |  |  |  |
Revenue recognized |  |  |  |  | $ 7,000,000 |  |  |  |  |  |
Revenue under collaborative research and development arrangements |  |  |  |  |  |  |  |  |  | 1,300,000 | $ 306,000
Deferred revenue |  |  |  | 1,200,000 |  |  |  |  |  | 1,200,000 |
Accounts receivable |  |  |  | 3,000,000 |  |  |  |  |  | 3,000,000 |
Roche |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |  |  |  |  |
Full payment |  |  |  |  |  | $ 8,500,000 |  |  |  |  |
Hoffman-La Roche |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |  |  |  |  |
Upfront payment received |  |  |  |  |  |  |  |  | $ 10,000,000 |  |
Previously deferred revenue recognized |  |  |  |  |  |  |  |  |  | 0 | 4,000,000
Defense Advanced Research Projects Agency (DARPA) |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  | 111,000 |  |  |  |  |  | 111,000 |
Accounts receivable |  |  |  | 2,000,000 |  |  |  |  |  | 2,000,000 |
Previously deferred revenue recognized |  |  |  |  |  |  |  |  |  |  | $ 5,000,000
Term |  |  |  |  |  |  |  | 2 years |  |  |
Base award |  |  |  |  |  |  |  | $ 19,600,000 |  |  |
Option award |  |  |  |  |  |  |  | 24,600,000 |  |  |
Second option award |  |  |  |  |  |  |  | $ 11,100,000 |  |  |
Funding received for research and development |  |  |  |  |  |  |  |  |  | $ 376,000 |
Foreign Tax Authority | ApolloBio |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |  |  |  |  |
Foreign corporate income taxes reducing upfront payment |  |  |  | 2,200,000 |  |  |  |  |  |  |
Foreign non-corporate income taxes reducing upfront payment |  |  |  | $ 1,400,000 |  |  |  |  |  |  |

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