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Condensed Consolidated Statements of Operations - USD ($)
 3 Months Ended
Mar. 31, 2018
Mar. 31, 2017
Revenues:    
Revenue under collaborative research and development arrangements $ 1,289,046 $ 4,288,586
Revenue under collaborative research and development arrangements with affiliated entities 148,008 233,330
Grants and miscellaneous revenue 92,590 5,240,233
Grants and miscellaneous revenue from affiliated entity 0 614,036
Total revenues 1,529,644 10,376,185
Operating expenses:    
Research and development 24,577,751 24,542,504
General and administrative 9,698,015 7,767,589
Total operating expenses 34,275,766 32,310,093
Loss from operations (32,746,122) (21,933,908)
Other income (expense):    
Interest and other income, net 312,523 340,341
Change in fair value of common stock warrants (127,841) 116,477
Gain (loss) on investment in affiliated entities 2,380,423 (1,608,817)
Net loss before provision for income taxes (30,181,017) (23,085,907)
Provision for income taxes (2,169,811) 0
Net loss $ (32,350,828) $ (23,085,907)
Net loss per share    
Basic (dollars per share) $ (0.36) $ (0.31)
Diluted (dollars per share) $ (0.36) $ (0.31)
Weighted average number of common shares outstanding    
Basic (shares) 90,451,791 74,152,609
Diluted (shares) 90,451,791 74,300,884