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Impact of Recently Issued Accounting Standards - Impact of Adopting ASU 2014-09 (Details) - USD ($) | 3 Months Ended | 6 Months Ended
--- | --- | ---
Jun. 30, 2018 | Jun. 30, 2017 | Jun. 30, 2018 | Jun. 30, 2017
Revenues [Abstract] |  |  |  |
Total revenues | $ 24,448,761 | $ 20,412,124 | $ 25,978,405 | $ 30,788,309
Operating Expenses [Abstract] |  |  |  |
Research and development | 22,462,620 | 23,878,751 | 47,040,371 | 48,421,255
General and administrative | 7,189,310 | 6,169,106 | 16,887,325 | 13,936,695
Total operating expenses | 29,651,930 | 30,047,857 | 63,927,696 | 62,357,950
Calculated under Revenue Guidance in Effect before Topic 606 |  |  |  |
Revenues [Abstract] |  |  |  |
Total revenues | 26,360,675 |  | 30,112,470 |
Operating Expenses [Abstract] |  |  |  |
Research and development | 24,374,534 |  | 51,174,436 |
General and administrative | 7,189,310 |  | 16,887,325 |
Total operating expenses | 31,563,844 |  | 68,061,761 |
Accounting Standards Update 2014-09 | Difference between Revenue Guidance in Effect before and after Topic 606 |  |  |  |
Revenues [Abstract] |  |  |  |
Total revenues | (1,911,914) |  | (4,134,065) |
Operating Expenses [Abstract] |  |  |  |
Research and development | (1,911,914) |  | (4,134,065) |
General and administrative | 0 |  | 0 |
Total operating expenses | (1,911,914) |  | (4,134,065) |
Revenue under collaborative research and development arrangements |  |  |  |
Revenues [Abstract] |  |  |  |
Total revenues | 24,385,852 | 16,358,316 | 25,674,898 | 20,646,902
Revenue under collaborative research and development arrangements | Calculated under Revenue Guidance in Effect before Topic 606 |  |  |  |
Revenues [Abstract] |  |  |  |
Total revenues | 24,385,852 |  | 25,674,898 |
Revenue under collaborative research and development arrangements | Accounting Standards Update 2014-09 | Difference between Revenue Guidance in Effect before and after Topic 606 |  |  |  |
Revenues [Abstract] |  |  |  |
Total revenues | 0 |  | 0 |
Revenue under collaborative research and development arrangements with affiliated entities |  |  |  |
Revenues [Abstract] |  |  |  |
Total revenues | 60,319 | 176,879 | 208,327 | 410,209
Revenue under collaborative research and development arrangements with affiliated entities | Calculated under Revenue Guidance in Effect before Topic 606 |  |  |  |
Revenues [Abstract] |  |  |  |
Total revenues | 60,319 |  | 208,327 |
Revenue under collaborative research and development arrangements with affiliated entities | Accounting Standards Update 2014-09 | Difference between Revenue Guidance in Effect before and after Topic 606 |  |  |  |
Revenues [Abstract] |  |  |  |
Total revenues | 0 |  | 0 |
Grants and miscellaneous revenue |  |  |  |
Revenues [Abstract] |  |  |  |
Total revenues | 2,590 | 2,797,647 | 95,180 | 8,037,880
Grants and miscellaneous revenue | Calculated under Revenue Guidance in Effect before Topic 606 |  |  |  |
Revenues [Abstract] |  |  |  |
Total revenues | 1,459,684 |  | 2,016,674 |
Grants and miscellaneous revenue | Accounting Standards Update 2014-09 | Difference between Revenue Guidance in Effect before and after Topic 606 |  |  |  |
Revenues [Abstract] |  |  |  |
Total revenues | (1,457,094) |  | (1,921,494) |
Grants and miscellaneous revenue from affiliated entity |  |  |  |
Revenues [Abstract] |  |  |  |
Total revenues | 0 | $ 1,079,282 | 0 | $ 1,693,318
Grants and miscellaneous revenue from affiliated entity | Calculated under Revenue Guidance in Effect before Topic 606 |  |  |  |
Revenues [Abstract] |  |  |  |
Total revenues | 454,820 |  | 2,212,571 |
Grants and miscellaneous revenue from affiliated entity | Accounting Standards Update 2014-09 | Difference between Revenue Guidance in Effect before and after Topic 606 |  |  |  |
Revenues [Abstract] |  |  |  |
Total revenues | $ (454,820) |  | $ (2,212,571) |

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