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Condensed Consolidated Statements of Operations - USD ($)
 3 Months Ended 6 Months Ended
Jun. 30, 2018
Jun. 30, 2017
Jun. 30, 2018
Jun. 30, 2017
Revenues:        
Total revenues $ 24,448,761 $ 20,412,124 $ 25,978,405 $ 30,788,309
Operating expenses:        
Research and development 22,462,620 23,878,751 47,040,371 48,421,255
General and administrative 7,189,310 6,169,106 16,887,325 13,936,695
Total operating expenses 29,651,930 30,047,857 63,927,696 62,357,950
Loss from operations (5,203,169) (9,635,733) (37,949,291) (31,569,641)
Other income (expense):        
Interest and other income, net 396,681 300,021 709,204 640,362
Change in fair value of common stock warrants 259,971 (312,500) 132,130 (196,023)
Gain (loss) on investment in affiliated entities (2,092,608) 169,096 287,815 (1,439,721)
Net loss before provision for income taxes (6,639,125) (9,479,116) (36,820,142) (32,565,023)
Provision for income taxes 0 (2,169,811) 0
Net loss $ (6,639,125) $ (9,479,116) $ (38,989,953) $ (32,565,023)
Net loss per share        
Basic (dollars per share) $ (0.07) $ (0.13) $ (0.43) $ (0.44)
Diluted (dollars per share) $ (0.08) $ (0.13) $ (0.43) $ (0.44)
Weighted average number of common shares outstanding        
Basic (shares) 91,153,776 75,409,702 90,804,722 74,783,791
Diluted (shares) 91,241,660 75,409,702 90,890,792 74,783,791
Revenue under collaborative research and development arrangements        
Revenues:        
Total revenues $ 24,385,852 $ 16,358,316 $ 25,674,898 $ 20,646,902
Revenue under collaborative research and development arrangements with affiliated entities        
Revenues:        
Total revenues 60,319 176,879 208,327 410,209
Grants and miscellaneous revenue        
Revenues:        
Total revenues 2,590 2,797,647 95,180 8,037,880
Grants and miscellaneous revenue from affiliated entity        
Revenues:        
Total revenues $ 0 $ 1,079,282 $ 0 $ 1,693,318