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Collaborative Agreements (Details)					1 Months Ended						3 Months Ended		9 Months Ended
	Apr. 11, 2018 USD ($)	Dec. 31, 2017 USD ($)	Dec. 29, 2017	Aug. 07, 2015 product	Mar. 31, 2018 USD ($)	Dec. 31, 2017 USD ($)	Feb. 28, 2017 USD ($)	Sep. 30, 2015 USD ($)	Apr. 30, 2015 USD ($)	Sep. 30, 2013 USD ($)	Sep. 30, 2018 USD ($)	Sep. 30, 2017 USD ($)	Sep. 30, 2018 USD ($)	Sep. 30, 2017 USD ($)
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues											$ 2,000,868	$ 2,644,543	$ 27,979,273	$ 33,432,852
Revenue recognized											316,000		1,300,000
Deferred grant funding		$ 0				$ 0					1,297,416		1,297,416
ApolloBio
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues											0		23,000,000
Deferred revenue		0				0					0		0
Revenue recognized													23,000,000
MedImmune
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues											1,800,000		4,500,000
Deferred revenue		1,145,500				1,145,500					295,533		295,533
Accounts receivable		1,693,530				1,693,530					2,352,323		2,352,323
Revenue recognized													1,582,167
Collaborative Arrangement, Product | ApolloBio
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Territory expansion option period			3 years
Proceeds received in upfront payment					$ 19,400,000
Upfront payment received					23,000,000						23,000,000
Advisory fees					960,000
Additional revenue to be achieved													$ 20,000,000
Royalty period													10 years
Period from effective date for termination													1 year
Number of days written notice before termination													90 days
Collaborative Arrangement, Product | MedImmune
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Upfront payment received		$ 27,500,000						$ 27,500,000
Anticipated development and regulatory event based payment receivable milestones													$ 355,000,000
Anticipated commercial event based payment receivable milestones													345,000,000
Number of additional products to be developed | product				2
Milestone payment, revenue recognized						$ 7,000,000
Deferred revenue											296,000		296,000
Accounts receivable											2,400,000		2,400,000
Revenue under collaborative research and development arrangements												344,000		14,900,000
Collaborative Arrangement, Product | Hoffman-La Roche
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Upfront payment received										$ 10,000,000
Full payment							$ 8,500,000
Revenue recognized											0	0	0	6,100,000
Collaborative Arrangement, Product | Coalition for Epidemic Preparedness Innovations
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Collaborative agreement, funding to be received	$ 56,000,000
Collaborative agreement, period to receive funding for research and development	5 years
Funding received for research and development											1,200,000		2,000,000
Deferred grant funding											1,300,000		1,300,000
Collaborative Arrangement, Product | Defense Advanced Research Projects Agency (DARPA)
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Accounts receivable											379,000		379,000
Funding received for research and development											140,000	$ 1,400,000	1,000,000	$ 9,100,000
Deferred grant funding											99,000		99,000
Base award									$ 19,600,000
Option award									$ 24,600,000
Total funding received for research and development											$ 43,700,000		$ 43,700,000
Foreign Tax Authority | Collaborative Arrangement, Product | ApolloBio
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Foreign corporate income taxes reducing upfront payment					2,200,000
Foreign non-corporate income taxes reducing upfront payment					$ 1,400,000

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