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Summary of Significant Accounting Policies - Impact of Adoption of ASU 2014-09 (Details) - USD ($)	3 Months Ended								12 Months Ended
	Dec. 31, 2018	Sep. 30, 2018	Jun. 30, 2018	Mar. 31, 2018	Dec. 31, 2017	Sep. 30, 2017	Jun. 30, 2017	Mar. 31, 2017	Dec. 31, 2018	Dec. 31, 2017	Dec. 31, 2016
Revenues:
Total revenues	$ 2,502,624	$ 2,000,868	$ 24,448,761	$ 1,529,644	$ 8,787,234	$ 2,644,543	$ 20,412,124	$ 10,376,185	$ 30,481,897	$ 42,220,086	$ 35,368,361
Operating expenses:
Research and development	26,365,647	21,851,858	22,462,620	24,577,751	24,641,124	25,510,239	23,878,751	24,542,504	95,257,876	98,572,618	88,712,035
General and administrative	5,636,141	6,791,693	7,189,310	9,698,015	8,033,899	6,319,775	6,169,106	7,767,589	29,315,159	28,290,369	23,892,263
Total operating expenses	32,001,788	28,643,551	29,651,930	34,275,766	31,675,023	31,830,014	30,047,857	32,310,093	124,573,035	125,862,987	111,604,298
Calculated under Revenue Guidance in Effect before Topic 606
Revenues:
Total revenues									39,973,602
Operating expenses:
Research and development									104,749,581
General and administrative									29,315,159
Total operating expenses									134,064,740
Revenue under collaborative research and development arrangements
Revenues:
Total revenues	2,372,600	1,813,287	24,385,852	1,289,046	7,409,214	351,272	16,358,316	4,288,586	29,860,785	28,407,388	6,490,747
Revenue under collaborative research and development arrangements | Calculated under Revenue Guidance in Effect before Topic 606
Revenues:
Total revenues									29,860,785
Revenue under collaborative research and development arrangements with affiliated entity
Revenues:
Total revenues	56,207	184,990	60,319	148,008	226,486	129,133	176,879	233,330	449,524	765,828	1,400,594
Revenue under collaborative research and development arrangements with affiliated entity | Calculated under Revenue Guidance in Effect before Topic 606
Revenues:
Total revenues									449,524
Grants and miscellaneous revenue
Revenues:
Total revenues	$ 73,817	$ 2,591	$ 2,590	$ 92,590	980,443	1,456,216	2,797,647	5,240,233	171,588	10,474,539	27,136,457
Grants and miscellaneous revenue | Calculated under Revenue Guidance in Effect before Topic 606
Revenues:
Total revenues									6,315,116
Grants and miscellaneous revenue from affiliated entity
Revenues:
Total revenues					$ 171,091	$ 707,922	$ 1,079,282	$ 614,036	0	$ 2,572,331	$ 340,563
Grants and miscellaneous revenue from affiliated entity | Calculated under Revenue Guidance in Effect before Topic 606
Revenues:
Total revenues									3,348,177
Accounting Standards Update 2014-09 | Difference between Revenue Guidance in Effect before and after Topic 606
Revenues:
Total revenues									(9,491,705)
Operating expenses:
Research and development									(9,491,705)
General and administrative									0
Total operating expenses									(9,491,705)
Accounting Standards Update 2014-09 | Revenue under collaborative research and development arrangements | Difference between Revenue Guidance in Effect before and after Topic 606
Revenues:
Total revenues									0
Accounting Standards Update 2014-09 | Revenue under collaborative research and development arrangements with affiliated entity | Difference between Revenue Guidance in Effect before and after Topic 606
Revenues:
Total revenues									0
Accounting Standards Update 2014-09 | Grants and miscellaneous revenue | Difference between Revenue Guidance in Effect before and after Topic 606
Revenues:
Total revenues									(6,143,528)
Accounting Standards Update 2014-09 | Grants and miscellaneous revenue from affiliated entity | Difference between Revenue Guidance in Effect before and after Topic 606
Revenues:
Total revenues									$ (3,348,177)

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