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Collaborative Agreements (Details)					1 Months Ended			3 Months Ended		12 Months Ended
	Apr. 11, 2018 USD ($)	Dec. 31, 2017 USD ($)	Dec. 29, 2017	Aug. 07, 2015 product	Mar. 31, 2018 USD ($)	Sep. 30, 2015 USD ($)	Apr. 30, 2015 USD ($)	Mar. 31, 2019 USD ($)	Mar. 31, 2018 USD ($)	Dec. 31, 2018 USD ($)
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues								$ 2,829,905	$ 1,529,644
Deferred grant funding								1,279,005		$ 4,165,848
AstraZeneca
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues								2,700,000	1,300,000
Collaborative Arrangement, Product | ApolloBio
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Territory expansion option period			3 years
Proceeds received in upfront payment					$ 19,400,000
Upfront payment received					23,000,000			23,000,000		$ 23,000,000
Advisory fees					960,000				960,000
Additional revenue to be achieved								$ 20,000,000
Royalty period								10 years
Period from effective date for termination								1 year
Number of days written notice before termination								90 days
Collaborative Arrangement, Product | AstraZeneca
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Upfront payment received		$ 27,500,000				$ 27,500,000
Anticipated development and regulatory event based payment receivable milestones								$ 355,000,000
Anticipated commercial event based payment receivable milestones								345,000,000
Number of additional products to be developed | product				2
Deferred revenue								169,000
Accounts receivable								2,600,000
Collaborative Arrangement, Product | Coalition for Epidemic Preparedness Innovations
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Collaborative agreement, funding to be received	$ 56,000,000
Collaborative agreement, period to receive funding for research and development	5 years
Funding received for research and development								1,700,000
Deferred grant funding								424,000
Collaborative Arrangement, Product | Defense Advanced Research Projects Agency (DARPA)
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Funding received for research and development								96,000	$ 376,000
Base award							$ 19,600,000
Option award							$ 24,600,000
Total funding received for research and development								$ 44,200,000
Foreign Tax Authority | Collaborative Arrangement, Product | ApolloBio
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Foreign corporate income taxes reducing upfront payment					2,200,000
Foreign non-corporate income taxes reducing upfront payment					$ 1,400,000

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