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Collaborative Agreements (Details) |  |  |  |  | 1 Months Ended | 3 Months Ended |
--- | --- | --- | --- | --- | --- | --- | ---
Apr. 11, 2018 USD ($) | Dec. 31, 2017 USD ($) | Dec. 29, 2017 | Aug. 07, 2015 product | Mar. 31, 2020 USD ($) | Jan. 31, 2020 USD ($) | Aug. 30, 2019 USD ($) | Oct. 31, 2018 USD ($) | Mar. 31, 2018 USD ($) | Sep. 30, 2015 USD ($) | Mar. 31, 2020 USD ($) | Mar. 31, 2019 USD ($) | Jun. 30, 2018 USD ($) | Dec. 31, 2019 USD ($)
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  | $ 1,327,274 | $ 2,829,905 |  |
Grant funding liability |  |  |  |  | $ 13,091,424 |  |  |  |  |  | 13,091,424 |  |  | $ 6,065,212
AstraZeneca |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  | 72,000 | 2,700,000 |  |
Collaborative Arrangement, Product | ApolloBio |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Territory expansion option period |  |  | 3 years |  |  |  |  |  |  |  |  |  |  |
Proceeds received in upfront payment |  |  |  |  |  |  |  |  | $ 19,400,000 |  |  |  |  |
Upfront payment received |  |  |  |  |  |  |  |  | 23,000,000 |  | 23,000,000 |  | $ 23,000,000 |
Advisory fees |  |  |  |  |  |  |  |  | 960,000 |  |  |  |  |
Additional revenue to be achieved |  |  |  |  |  |  |  |  |  |  | $ 20,000,000 |  |  |
Royalty period |  |  |  |  |  |  |  |  |  |  | 10 years |  |  |
Period from effective date for termination |  |  |  |  |  |  |  |  |  |  | 1 year |  |  |
Number of days written notice before termination |  |  |  |  |  |  |  |  |  |  | 90 days |  |  |
Collaborative Arrangement, Product | AstraZeneca |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Upfront payment received |  | $ 27,500,000 |  |  |  |  |  |  |  | $ 27,500,000 |  |  |  |
Anticipated development and regulatory event based payment receivable milestones |  |  |  |  |  |  |  |  |  |  | $ 125,000,000 |  |  |
Anticipated commercial event based payment receivable milestones |  |  |  |  |  |  |  |  |  |  | 115,000,000 |  |  |
Number of additional products to be developed | product |  |  |  | 2 |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  |  | 62,000 |  |  |  |  |  | 62,000 |  |  |
Foreign Tax Authority | Collaborative Arrangement, Product | ApolloBio |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Foreign corporate income taxes reducing upfront payment |  |  |  |  |  |  |  |  | 2,200,000 |  |  |  |  |
Foreign non-corporate income taxes reducing upfront payment |  |  |  |  |  |  |  |  | $ 1,400,000 |  |  |  |  |
Lassa Fever And MERS Vaccine | Collaborative Arrangement, Product | Coalition for Epidemic Preparedness Innovations |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative agreement, funding to be received | $ 56,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative agreement, period to receive funding for research and development | 5 years |  |  |  |  |  |  |  |  |  |  |  |  |
Funding received for research and development |  |  |  |  |  |  |  |  |  |  | 987,000 | 1,700,000 |  |
Grant funding liability |  |  |  |  | 4,400,000 |  |  |  |  |  | 4,400,000 |  |  |
SARS-CoV-2/COVID-19 Vaccine | Collaborative Arrangement, Product | Coalition for Epidemic Preparedness Innovations |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative agreement, funding to be received |  |  |  |  |  | $ 9,000,000 |  |  |  |  |  |  |  |
Funding received for research and development |  |  |  |  |  |  |  |  |  |  | 2,400,000 |  |  |
Grant funding liability |  |  |  |  | 2,900,000 |  |  |  |  |  | 2,900,000 |  |  |
DNA-Encoded Monoclonal Antibody Technology | Collaborative Arrangement, Product | Bill And Melinda Gates Foundation |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Funding received for research and development |  |  |  |  |  |  | $ 1,100,000 | $ 2,200,000 |  |  | 134,000 | $ 1,100,000 |  |
Grant funding liability |  |  |  |  | 892,000 |  |  |  |  |  | 892,000 |  |  |
CELLECTRA 3PSP Proprietary Smart Device | Collaborative Arrangement, Product | Bill And Melinda Gates Foundation |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Funding received for research and development |  |  |  |  | 5,000,000 |  |  |  |  |  | 64,000 |  |  |
Grant funding liability |  |  |  |  | $ 4,900,000 |  |  |  |  |  | $ 4,900,000 |  |  |

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