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Consolidated Statements of Stockholders' Equity - USD ($) | Total | 6.50% Convertible Senior Notes Due 2024 | August 2019 Bonds | Cumulative Effect, Period of Adoption, Adjustment | [1] | Preferred stock | Common stock | Common stock 6.50% Convertible Senior Notes Due 2024 | Common stock August 2019 Bonds | Additional paid-in capital | Additional paid-in capital 6.50% Convertible Senior Notes Due 2024 | Additional paid-in capital August 2019 Bonds | Accumulated deficit | Accumulated deficit Cumulative Effect, Period of Adoption, Adjustment | [1] | Accumulated other comprehensive income (loss) | Accumulated other comprehensive income (loss) Cumulative Effect, Period of Adoption, Adjustment | [1] | Non- controlling interest
--- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | ---
Beginning balance (in shares) at Dec. 31, 2017 |  |  |  |  | 23 | 90,357,644 |  |  |  |  |  |  |  |  |  |
Beginning balance at Dec. 31, 2017 | $ 142,488,809 |  |  | $ 0 | $ 0 | $ 90,358 |  |  | $ 665,775,504 |  |  | $ (523,356,317) | $ 231,366 | $ (117,005) | $ (231,366) | $ 96,269
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock for cash (in shares) |  |  |  |  |  | 5,669,025 |  |  |  |  |  |  |  |  |  |
Issuance of common stock for cash | 29,227,776 |  |  |  |  | $ 5,669 |  |  | 29,222,107 |  |  |  |  |  |  |
Exercise of stock options and warrants for cash and vesting of RSUs, net of tax payments (in shares) |  |  |  |  |  | 1,199,141 |  |  |  |  |  |  |  |  |  |
Exercise of stock options and warrants for cash and vesting of RSUs, net of tax payments | 1,809,526 |  |  |  |  | $ 1,199 |  |  | 1,808,327 |  |  |  |  |  |  |
Stock-based compensation | 10,654,622 |  |  |  |  |  |  |  | 10,988,277 |  |  | (333,655) |  |  |  |
Net loss attributable to common stockholders | (96,967,830) |  |  |  |  |  |  |  |  |  |  | (96,967,830) |  |  |  |
Unrealized gain (loss) on short-term investments, net of tax | (180,496) |  |  |  |  |  |  |  |  |  |  |  |  | (180,496) |  |
Foreign currency translation | 0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Ending balance (in shares) at Dec. 31, 2018 |  |  |  |  | 23 | 97,225,810 |  |  |  |  |  |  |  |  |  |
Ending balance at Dec. 31, 2018 | 87,032,407 |  |  |  | $ 0 | $ 97,226 |  |  | 707,794,215 |  |  | (620,426,436) |  | (528,867) |  | 96,269
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock for cash (in shares) |  |  |  |  |  | 3,340,678 |  |  |  |  |  |  |  |  |  |
Issuance of common stock for cash | 9,089,009 |  |  |  |  | $ 3,340 |  |  | 9,085,669 |  |  |  |  |  |  |
Exercise of stock options and warrants for cash and vesting of RSUs, net of tax payments (in shares) |  |  |  |  |  | 794,546 |  |  |  |  |  |  |  |  |  |
Exercise of stock options and warrants for cash and vesting of RSUs, net of tax payments | (780,405) |  |  |  |  | $ 795 |  |  | (781,200) |  |  |  |  |  |  |
Equity component of issuance of convertible notes | 15,752,698 |  |  |  |  |  |  |  | 15,752,698 |  |  |  |  |  |  |
Stock-based compensation | 10,901,320 |  |  |  |  |  |  |  | 10,795,403 |  |  |  |  |  |  | 105,917
Acquisition of non-controlling interest in Geneos, net | 2,960,131 |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 2,960,131
Net loss attributable to common stockholders | (120,551,777) |  |  |  |  |  |  |  |  |  |  | (119,359,219) |  |  |  | (1,192,558)
Unrealized gain (loss) on short-term investments, net of tax | 1,001,475 |  |  |  |  |  |  |  |  |  |  |  |  | 1,001,475 |  |
Foreign currency translation | 0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Ending balance (in shares) at Dec. 31, 2019 |  |  |  |  | 23 | 101,361,034 |  |  |  |  |  |  |  |  |  |
Ending balance at Dec. 31, 2019 | 5,404,858 |  |  |  | $ 0 | $ 101,361 |  |  | 742,646,785 |  |  | (739,785,655) |  | 472,608 |  | 1,969,759
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock for cash (in shares) |  |  |  |  |  | 66,064,886 |  |  |  |  |  |  |  |  |  |
Issuance of common stock for cash | 454,486,400 |  |  |  |  | $ 66,065 |  |  | 454,420,335 |  |  |  |  |  |  |
Conversion of preferred stock/senior notes to common stock (in shares) |  |  |  |  | (14) | 5,147 | 11,535,660 | 4,962,364 |  |  |  |  |  |  |  |
Conversion of preferred stock/senior notes to common stock | 0 | $ 43,694,386 | $ 102,671,310 |  |  | $ 5 | $ 11,536 | $ 4,961 | (5) | $ 43,682,850 | $ 102,666,349 |  |  |  |  |
Exercise of stock options and warrants for cash and vesting of RSUs, net of tax payments (in shares) |  |  |  |  |  | 2,922,402 |  |  |  |  |  |  |  |  |  |
Exercise of stock options and warrants for cash and vesting of RSUs, net of tax payments | 8,241,624 |  |  |  |  | $ 2,923 |  |  | 8,238,701 |  |  |  |  |  |  |
Stock-based compensation | 15,647,523 |  |  |  |  |  |  |  | 15,655,585 |  |  |  |  |  |  | (8,062)
Acquisition of non-controlling interest in Geneos, net | 2,379,969 |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 2,379,969
Deconsolidation of Geneos | (3,181,640) |  |  |  |  |  |  |  |  |  |  |  |  |  |  | (3,181,640)
Net loss attributable to common stockholders | (167,474,914) |  |  |  |  |  |  |  |  |  |  | (166,411,157) |  |  |  | (1,063,757)
Dissolution of majority-owned subsidiary VGX Animal Health, Inc. | 0 |  |  |  |  |  |  |  | 96,269 |  |  |  |  |  |  | (96,269)
Unrealized gain (loss) on short-term investments, net of tax | (755,963) |  |  |  |  |  |  |  |  |  |  |  |  | (755,963) |  |
Foreign currency translation | 27,205 |  |  |  |  |  |  |  |  |  |  |  |  | 27,205 |  |
Ending balance (in shares) at Dec. 31, 2020 |  |  |  |  | 9 | 186,851,493 |  |  |  |  |  |  |  |  |  |
Ending balance at Dec. 31, 2020 | $ 461,140,758 |  |  |  | $ 0 | $ 186,851 |  |  | $ 1,367,406,869 |  |  | $ (906,196,812) |  | $ (256,150) |  | $ 0
[1] | Upon adoption of ASU 2016-01 on January 1, 2018, the Company began to measure and record equity investments, except those accounted for under the equity method of accounting that have a readily determinable fair value, at fair value and began to recognize the changes in fair value in its consolidated statements of operations, instead of recognizing unrealized gains and losses through accumulated other comprehensive income (loss), as done under the previous guidance. The Company recorded a $231,000 cumulative effect adjustment to reclassify the cumulative unrealized gain, net of tax effect, from its investment in an affiliated entity, PLS, from accumulated other comprehensive loss to accumulated deficit.

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