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Consolidated Statements of Stockholders' Equity - USD ($) | Total | 6.50% Convertible Senior Notes Due 2024 | August 2019 Bonds | Preferred stock | Common stock | Common stock 6.50% Convertible Senior Notes Due 2024 | Common stock August 2019 Bonds | Additional paid-in capital | Additional paid-in capital 6.50% Convertible Senior Notes Due 2024 | Additional paid-in capital August 2019 Bonds | Accumulated deficit | Accumulated other comprehensive income (loss) | Non- controlling interest
--- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | ---
Beginning balance (in shares) at Dec. 31, 2018 |  |  |  | 23 | 97,225,810 |  |  |  |  |  |  |  |
Beginning balance at Dec. 31, 2018 | $ 87,032,407 |  |  | $ 0 | $ 97,226 |  |  | $ 707,794,215 |  |  | $ (620,426,436) | $ (528,867) | $ 96,269
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock for cash, net of capital raising related expenses (in shares) |  |  |  |  | 3,340,678 |  |  |  |  |  |  |  |
Issuance of common stock for cash, net of capital raising related expenses | 9,089,009 |  |  |  | $ 3,340 |  |  | 9,085,669 |  |  |  |  |
Exercise of stock options and warrants for cash and vesting of RSUs, net of tax payments (in shares) |  |  |  |  | 794,546 |  |  |  |  |  |  |  |
Exercise of stock options and warrants for cash and vesting of RSUs, net of tax payments | (780,405) |  |  |  | $ 795 |  |  | (781,200) |  |  |  |  |
Equity component of issuance of convertible notes | 15,752,698 |  |  |  |  |  |  | 15,752,698 |  |  |  |  |
Stock-based compensation | 10,901,320 |  |  |  |  |  |  | 10,795,403 |  |  |  |  | 105,917
Acquisition of non-controlling interest in Geneos, net | 2,960,131 |  |  |  |  |  |  |  |  |  |  |  | 2,960,131
Net loss attributable to common stockholders | (120,551,777) |  |  |  |  |  |  |  |  |  | (119,359,219) |  | (1,192,558)
Unrealized gain/loss on short-term investments, net of tax | 1,001,475 |  |  |  |  |  |  |  |  |  |  | 1,001,475 |
Foreign currency translation | 0 |  |  |  |  |  |  |  |  |  |  |  |
Ending balance (in shares) at Dec. 31, 2019 |  |  |  | 23 | 101,361,034 |  |  |  |  |  |  |  |
Ending balance at Dec. 31, 2019 | 5,404,858 |  |  | $ 0 | $ 101,361 |  |  | 742,646,785 |  |  | (739,785,655) | 472,608 | 1,969,759
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock for cash, net of capital raising related expenses (in shares) |  |  |  |  | 66,064,886 |  |  |  |  |  |  |  |
Issuance of common stock for cash, net of capital raising related expenses | 454,486,400 |  |  |  | $ 66,065 |  |  | 454,420,335 |  |  |  |  |
Conversion of preferred stock/senior notes to common stock (in shares) |  |  |  | (14) | 5,147 | 11,535,660 | 4,962,364 |  |  |  |  |  |
Conversion of preferred stock/senior notes to common stock | 0 | $ 43,694,386 | $ 102,671,310 |  | $ 5 | $ 11,536 | $ 4,961 | (5) | $ 43,682,850 | $ 102,666,349 |  |  |
Exercise of stock options and warrants for cash and vesting of RSUs, net of tax payments (in shares) |  |  |  |  | 2,922,402 |  |  |  |  |  |  |  |
Exercise of stock options and warrants for cash and vesting of RSUs, net of tax payments | 8,241,624 |  |  |  | $ 2,923 |  |  | 8,238,701 |  |  |  |  |
Stock-based compensation | 15,647,523 |  |  |  |  |  |  | 15,655,585 |  |  |  |  | (8,062)
Acquisition of non-controlling interest in Geneos, net | 2,379,969 |  |  |  |  |  |  |  |  |  |  |  | 2,379,969
Deconsolidation of Geneos | (3,181,640) |  |  |  |  |  |  |  |  |  |  |  | (3,181,640)
Net loss attributable to common stockholders | (167,474,914) |  |  |  |  |  |  |  |  |  | (166,411,157) |  | (1,063,757)
Dissolution of majority-owned subsidiary VGX Animal Health, Inc. | 0 |  |  |  |  |  |  | 96,269 |  |  |  |  | (96,269)
Unrealized gain/loss on short-term investments, net of tax | (755,963) |  |  |  |  |  |  |  |  |  |  | (755,963) |
Foreign currency translation | 27,205 |  |  |  |  |  |  |  |  |  |  | 27,205 |
Ending balance (in shares) at Dec. 31, 2020 |  |  |  | 9 | 186,851,493 |  |  |  |  |  |  |  |
Ending balance at Dec. 31, 2020 | 461,140,758 |  |  | $ 0 | $ 186,851 |  |  | 1,367,406,869 |  |  | (906,196,812) | (256,150) | 0
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock for cash, net of capital raising related expenses (in shares) |  |  |  |  | 27,310,341 |  |  |  |  |  |  |  |
Issuance of common stock for cash, net of capital raising related expenses | 209,441,410 |  |  |  | $ 27,310 |  |  | 209,414,100 |  |  |  |  |
Conversion of preferred stock/senior notes to common stock (in shares) |  |  |  |  |  |  | 1,009,450 |  |  |  |  |  |
Conversion of preferred stock/senior notes to common stock |  |  | $ 4,377,892 |  |  |  | $ 1,009 |  |  | $ 4,376,883 |  |  |
Exercise of stock options and warrants for cash and vesting of RSUs, net of tax payments (in shares) |  |  |  |  | 2,211,603 |  |  |  |  |  |  |  |
Exercise of stock options and warrants for cash and vesting of RSUs, net of tax payments | 2,057,393 |  |  |  | $ 2,212 |  |  | 2,055,181 |  |  |  |  |
Stock-based compensation | 26,336,764 |  |  |  |  |  |  | 26,336,764 |  |  |  |  | 0
Net loss attributable to common stockholders | (303,658,710) |  |  |  |  |  |  |  |  |  | (303,658,710) |  | 0
Unrealized gain/loss on short-term investments, net of tax | 4,048 |  |  |  |  |  |  |  |  |  |  | 4,048 |
Foreign currency translation | (30,134) |  |  |  |  |  |  |  |  |  |  | (30,134) |
Ending balance (in shares) at Dec. 31, 2021 |  |  |  | 9 | 217,382,887 |  |  |  |  |  |  |  |
Ending balance at Dec. 31, 2021 | $ 399,669,421 |  |  | $ 0 | $ 217,382 |  |  | $ 1,609,589,797 |  |  | $ (1,209,855,522) | $ (282,236) | $ 0

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