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Related Party Transactions (Details) - USD ($) $ in Thousands | 1 Months Ended | 12 Months Ended
--- | --- | ---
Nov. 30, 2016 | Mar. 31, 2016 | Dec. 31, 2021 | Dec. 31, 2020
Related Party Transaction [Line Items] |  |  |  |
Agreement amended amount |  |  | $ 13,500 |
Awarded option amount |  |  |  | $ 6,000
Awarded exercised amount |  |  | 3,300 |
Director |  |  |  |
Related Party Transaction [Line Items] |  |  |  |
Grant proceeds received |  |  | $ 3,000 | $ 1,900
Available-for-sale Securities | Plumbline Life Sciences |  |  |  |
Related Party Transaction [Line Items] |  |  |  |
Percentage of ownership |  |  | 18.90% | 19.70%
PLS (affiliated entity) |  |  |  |
Related Party Transaction [Line Items] |  |  |  |
Number of shares owned (in shares) |  |  | 597,808 |
Revenue from related parties |  |  | $ 245 | $ 1,400
Accounts receivable |  |  | 25 | 67
The Wistar Institute |  |  |  |
Related Party Transaction [Line Items] |  |  |  |
Accounts receivable |  |  | 2,600 | 425
Expenses to reimburse |  | $ 3,100 |  |
Term |  | 5 years |  |
Proceeds from sub-grant | $ 6,100 |  |  |
Awarded amount |  |  |  | 10,700
Operating expenses related to affiliated entity |  |  | 2,900 | 2,300
Due to related parties |  |  | 548 | 643
Deferred grant funding from affiliate |  |  | 37 |
Prepaid expense |  |  |  | $ 261
Director |  |  |  |
Related Party Transaction [Line Items] |  |  |  |
Expenses to reimburse |  |  | $ 1,900 |
Term |  |  | 2 years |

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