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Collaborative Agreements (Details) - USD ($)
 1 Months Ended 3 Months Ended 12 Months Ended
Apr. 11, 2018
Dec. 31, 2017
Jan. 31, 2021
Jun. 30, 2020
Apr. 30, 2020
Mar. 31, 2020
Jan. 31, 2020
Aug. 30, 2019
Mar. 31, 2019
Dec. 31, 2018
Oct. 31, 2018
Mar. 31, 2018
Sep. 30, 2015
Jun. 30, 2018
Dec. 31, 2021
Dec. 31, 2020
Dec. 31, 2019
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Total revenues                             $ 1,774,758 $ 7,411,220 $ 4,111,930
Deferred grant funding                             4,559,721 7,474,310  
Revenue under collaborative research and development arrangements                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Total revenues                             902,260 5,170,586 3,636,945
Advaccine                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Total revenues                             0 5,000,000 $ 0
Advaccine | Customer Concentration Risk                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Accounts receivable                               7,100,000  
Advaccine | Collaborative Arrangement                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Upfront payment received     $ 3,000,000                       4,500,000    
Additional revenue to be achieved                             206,000,000    
Collaboration agreement, payment earned                             $ 2,000,000 2,000,000  
Obligation period to pay royalties                             10 years    
Cost, Maintenance                             $ 1,500,000    
Annual Maintenance Period                             5 years    
Total revenues                             $ 5,000,000    
Agreement, number of days written notice before termination                             18 months    
Advaccine | Collaborative Arrangement | Revenue under collaborative research and development arrangements                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Total revenues                               5,000,000  
ApolloBio | Collaborative Arrangement                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Upfront payment received                       $ 23,000,000   $ 23,000,000 $ 23,000,000    
Additional revenue to be achieved                             $ 20,000,000    
Obligation period to pay royalties                             10 years    
Proceeds from collaborative agreement                       19,400,000          
Advisory fees                 $ 960,000                
Agreement period from effective date for termination                             1 year    
Agreement, number of days written notice before termination                             90 days    
ApolloBio | Collaborative Arrangement | Foreign Tax Authority                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Collaborative agreement, corporate income tax                       2,200,000          
Collaborative agreement, foreign non-income Taxes                       $ 1,400,000          
AstraZeneca | Collaborative Arrangement                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Upfront payment received   $ 27,500,000                     $ 27,500,000        
Agreement, milestone payment, revenue recognized                 $ 2,000,000 $ 2,000,000              
Coalition for Epidemic Preparedness Innovations | Collaborative Arrangement                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Upfront payment received                             $ 10,000,000 6,400,000  
Collaborative agreement, funding to be received $ 56,000,000       $ 6,900,000                        
Collaborative agreement, period to receive funding for research and development 5 years                                
Deferred grant funding                             23,000    
Coalition for Epidemic Preparedness Innovations | Collaborative Arrangement | Lassa Fever And MERS Vaccine                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Accounts receivable                             1,900,000    
Coalition for Epidemic Preparedness Innovations | Collaborative Arrangement | COVID19 Vaccine                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Collaborative agreement, funding to be received             $ 9,000,000               6,900,000 10,000,000  
Deferred grant funding                             1,800,000    
Coalition for Epidemic Preparedness Innovations | Collaborative Arrangement | CELLECTRA 3PSP Proprietary Smart Device                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Collaborative agreement, funding to be received         5,000,000                        
Coalition for Epidemic Preparedness Innovations | Collaborative Arrangement | IN O4800                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Collaborative agreement, funding to be received         $ 1,300,000                        
Bill And Melinda Gates Foundation | Collaborative Arrangement | CELLECTRA 3PSP Proprietary Smart Device                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Collaborative agreement, funding to be received           $ 5,000,000                 893,000 4,100,000  
Bill And Melinda Gates Foundation | Collaborative Arrangement | D N A Encoded Monoclonal Antibody Technology                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Collaborative agreement, funding to be received               $ 1,100,000     $ 2,200,000       182,000 463,000  
Deferred grant funding                             384,000    
Department Of Defence | Collaborative Arrangement                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Collaborative agreement, funding to be received       $ 54,500,000                          
Collaborative Arrangement , Revenue from the Procurement Contract                             755,000,000    
Grant proceeds received                             27,100,000 $ 21,200,000  
Accounts receivable                             3,600,000    
Grants Receivable                             2,300,000    
Department Of Defence | Collaborative Arrangement | CELLECTRA 3PSP Proprietary Smart Device                                  
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                                  
Purchase price, procurement contract                             $ 10,700,000