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Income Taxes - Significant Components of Deferred Tax Assets and Liabilities (Details) - USD ($) $ in Thousands | Dec. 31, 2023 | Dec. 31, 2022
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Deferred tax assets: |  |
Capitalized research expense | $ 50,143 | $ 41,252
NOL carryforwards | 235,624 | 212,768
Research and development and other tax credits | 27,734 | 26,442
Deferred revenue | 22 | 538
Stock-based compensation | 3,683 | 3,945
Acquired intangibles | 907 | 559
Investment in affiliated entity | 1,406 | 1,569
Lease liability | 2,822 | 3,247
Fixed assets | 337 | 57
Other | 6,808 | 11,062
Deferred tax assets, gross | 329,486 | 301,439
Valuation allowance | (327,493) | (299,124)
Total deferred tax assets | 1,993 | 2,315
Deferred tax liabilities: |  |
Acquired intangibles | 0 | (199)
Right of use asset | (1,993) | (2,148)
Total deferred tax liabilities | (1,993) | (2,347)
Net deferred tax liabilities | $ 0 | $ (32)

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