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Consolidated Statements of Cash Flows - USD ($) | 12 Months Ended
--- | ---
Dec. 31, 2023 | Dec. 31, 2022 | Dec. 31, 2021
Cash flows from operating activities: |  |  |
Net loss | $ (135,117,352) | $ (279,818,065) | $ (303,658,710)
Adjustments to reconcile net loss to net cash used in operating activities: |  |  |
Depreciation | 2,621,649 | 3,656,713 | 3,040,096
Amortization of intangible assets | 145,417 | 496,494 | 520,415
Amortization of operating lease right-of-use assets | 736,472 | 1,342,819 | 1,170,270
Impairment of goodwill | 10,513,371 | 0 | 0
Impairment of intangible assets | 1,984,444 | 0 | 0
Deferred taxes | (32,046) | 0 | 0
Non-cash stock-based compensation | 11,072,486 | 22,555,419 | 26,336,764
Non-cash interest on senior convertible notes | 155,814 | 186,977 | 858,644
Amortization of (discounts) premiums on investments | (4,686,144) | (1,320,546) | 1,633,286
Realized loss on sales of short-term investments | 4,805,804 | 4,029,961 | 5,397
Gain on remeasurement of investment in Geneos | 0 | (165,215) | 0
Net loss on disposal of fixed assets | 317,997 | 1,074,830 | 0
(Gain) loss on equity investment in affiliated entity | (773,145) | 1,899,654 | 553,570
Share of net loss in Geneos | 0 | 2,165,213 | 434,387
Net unrealized (gain) loss on available-for-sale equity securities | (5,850,626) | 7,846,172 | 3,222,838
Unrealized transaction (gain) loss on foreign-currency denominated debt | 0 | 0 | (176,927)
Changes in operating assets and liabilities: |  |  |
Accounts receivable, including from affiliated entities | 9,332,988 | (3,706,172) | 11,031,705
Prepaid expenses and other current assets, including from affiliated entities | 39,020,611 | (5,336,525) | (6,343,632)
Other assets | 78,729 | 741,750 | 24,531,654
Accounts payable and accrued expenses, including due to affiliated entities | (45,989,061) | 32,606,581 | 26,140,970
Accrued clinical trial expenses | (8,228,691) | 267,807 | 375,921
Deferred revenue, including from affiliated entity | 0 | (85,989) | (39,853)
Operating lease right-of-use assets and liabilities, net | (2,020,971) | (2,603,956) | (2,329,394)
Grant funding liability, including from affiliated entity | (2,453,297) | (2,034,517) | (2,973,089)
Other liabilities | 0 | (14,826) | (42,837)
Net cash used in operating activities | (124,365,551) | (216,215,421) | (215,708,525)
Cash flows from investing activities: |  |  |
Purchases of investments | (203,475,052) | (248,528,843) | (348,953,236)
Proceeds from sale or maturity of investments | 284,932,562 | 361,083,850 | 174,839,758
Purchases of capital assets | (320,898) | (969,153) | (1,231,006)
Proceeds from sale of capital assets | 6,219,263 | 0 | 0
Investment in Geneos | 0 | (1,999,998) | 0
Net cash provided by (used in) investing activities | 87,355,875 | 109,585,856 | (175,344,484)
Cash flows from financing activities: |  |  |
Proceeds from issuance of common stock, net of issuance costs | 5,461,745 | 82,955,311 | 209,441,410
Proceeds from stock option exercises | 0 | 283,022 | 6,668,741
Taxes paid related to net share settlement of equity awards | (466,646) | (1,397,631) | (4,611,348)
Net cash provided by financing activities | 4,995,099 | 81,840,702 | 211,498,803
Effect of exchange rate changes on cash and cash equivalents | (3,920) | (25,556) | (30,134)
Decrease in cash and cash equivalents | (32,018,497) | (24,814,419) | (179,584,340)
Cash and cash equivalents, beginning of period | 46,329,359 | 71,143,778 | 250,728,118
Cash and cash equivalents, end of period | 14,310,862 | 46,329,359 | 71,143,778
Supplemental disclosure: |  |  |
Amounts accrued for purchases of fixed assets | 0 | 108,181 | 204,815
Interest paid | 1,066,975 | 1,066,975 | 1,077,803
Change in prepaid expenses and other current assets related to fixed assets | 0 | 6,071,000 | 7,709,337
Issuance of common stock as part of litigation settlement | $ 14,000,000 | $ 0 | $ 0

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