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Stock-Based Compensation - Narrative (Details) - USD ($)
3 Months Ended
Mar. 31, 2024
Mar. 31, 2023
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]    
Non-cash stock-based compensation $ 2,525,433 $ 3,809,003
Total unrecognized compensation cost related to unvested stock options $ 4,300,000  
Period over which total unrecognized compensation cost related to unvested stock options will be recognized (in years) 1 year 8 months 12 days  
Weighted average grant date fair value (in dollars per share) $ 6.77 $ 11.64
Performance-based restricted stock units    
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]    
Total unrecognized compensation cost related to unvested stock options $ 4,100,000  
Period over which total unrecognized compensation cost related to unvested stock options will be recognized (in years) 1 year 10 months 24 days  
Weighted average grant date fair value, restricted stock units (in dollars per share) $ 8.39 $ 14.40
Employees and Directors    
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]    
Non-cash stock-based compensation $ 2,400,000 $ 3,600,000
Employees and Directors | Research and Development Expense    
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]    
Allocated share-based compensation expense 1,000,000 1,500,000
Employees and Directors | General and Administrative Expense    
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]    
Allocated share-based compensation expense 1,400,000 2,100,000
Non Employee    
Share-based Compensation Arrangement by Share-based Payment Award [Line Items]    
Non-cash stock-based compensation $ 103,000 $ 220,000