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CONSOLIDATED STATEMENTS OF CONVERTIBLE PREFERRED SHARES AND STOCKHOLDERS' EQUITY - USD ($) $ in Thousands | Total | Series C Preferred Stock [Member] | Series D Preferred Stock [Member] | Pfizer Purchase Agreement [Member] | Series A, B, C and D Convertible Preferred Stock [Member] | Series A, B, C and D Convertible Preferred Stock [Member] Series C Preferred Stock [Member] | Series A, B, C and D Convertible Preferred Stock [Member] Series D Preferred Stock [Member] | Common Stock [Member] | Common Stock [Member] Pfizer Purchase Agreement [Member] | Additional Paid-in Capital [Member] | Additional Paid-in Capital [Member] Series C Preferred Stock [Member] | Additional Paid-in Capital [Member] Series D Preferred Stock [Member] | Additional Paid-in Capital [Member] Pfizer Purchase Agreement [Member] | Accumulated Deficit [Member] | Accumulated Other Comprehensive Income (Loss) [Member]
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Balances at Dec. 31, 2019 | $ 74,574 |  |  |  |  |  |  | $ 19 |  | $ 273,966 |  |  |  | $ (199,427) | $ 16
Balance, Shares at Dec. 31, 2019 |  |  |  |  | 2,720 |  |  | 19,190,695 |  |  |  |  |  |  |
Issuance of common stock upon the exercise of stock options | 2,189 |  |  |  |  |  |  |  |  | 2,189 |  |  |  |  |
Issuance of common stock upon the exercise of stock options, shares |  |  |  |  |  |  |  | 324,433 |  |  |  |  |  |  |
Issuance of common stock and preferred stock net of offering costs and issuance costs | 77,929 | $ 20,542 | $ 30,221 |  |  |  | $ 3 | $ 8 |  | 77,921 | $ 20,542 | $ 30,218 |  |  |
Issuance of common stock and preferred stock net of offering costs, shares |  |  |  |  |  | 2,287 | 3,215,000 | 8,025,119 |  |  |  |  |  |  |
Beneficial conversion feature of Series C preferred stock |  |  |  |  |  | $ (549) |  |  |  |  | 549 |  |  |  |
Deemed dividends related to immediate accretion of beneficial conversion feature of Series C preferred stock |  |  |  |  |  | $ 549 |  |  |  |  | $ (549) |  |  |  |
Conversion of convertible preferred stock to common stock |  |  |  |  |  |  |  | $ 2 |  | (2) |  |  |  |  |
Conversion of convertible preferred stock to common stock, shares |  |  |  |  | (1,720) |  |  | 1,720,000 |  |  |  |  |  |  |
Share-based compensation expense | 4,888 |  |  |  |  |  |  |  |  | 4,888 |  |  |  |  |
Unrealized gain (loss) on available-for-sale securities | (23) |  |  |  |  |  |  |  |  |  |  |  |  |  | (23)
Net loss | (78,280) |  |  |  |  |  |  |  |  |  |  |  |  | (78,280) |
Balances at Dec. 31, 2020 | 132,040 |  |  |  | $ 3 |  |  | $ 29 |  | 409,722 |  |  |  | (277,707) | (7)
Balance, shares at Dec. 31, 2020 |  |  |  |  | 3,218,287 |  |  | 29,260,247 |  |  |  |  |  |  |
Issuance of common stock upon the exercise of stock options | $ 1,450 |  |  |  |  |  |  |  |  | 1,450 |  |  |  |  |
Issuance of common stock upon the exercise of stock options, shares | 160,674 |  |  |  |  |  |  | 160,674 |  |  |  |  |  |  |
Issuance of common stock and preferred stock net of offering costs and issuance costs | $ 7,831 |  |  | $ 27,289 |  |  |  | $ 1 | $ 2 | 7,830 |  |  | $ 27,287 |  |
Issuance of common stock and preferred stock net of offering costs, shares |  |  | 3,215,000 |  |  |  |  | 475,469 | 2,362,348 |  |  |  |  |  |
Conversion of convertible preferred stock to common stock, shares |  |  |  |  | (135) |  |  | 135,000 |  |  |  |  |  |  |
Share-based compensation expense | 9,430 |  |  |  |  |  |  |  |  | 9,430 |  |  |  |  |
Unrealized gain (loss) on available-for-sale securities | 5 |  |  |  |  |  |  |  |  |  |  |  |  |  | 5
Net loss | (89,756) |  |  |  |  |  |  |  |  |  |  |  |  | (89,756) |
Balances at Dec. 31, 2021 | $ 88,289 |  |  |  | $ 3 |  |  | $ 32 |  | $ 455,719 |  |  |  | $ (367,463) | $ (2)
Balance, shares at Dec. 31, 2021 |  |  |  |  | 3,218,152 |  |  | 32,393,738 |  |  |  |  |  |  |

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