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Property and Equipment (Details) - USD ($) $ in Thousands | 12 Months Ended
--- | ---
Dec. 31, 2015 | Dec. 31, 2014 | Dec. 31, 2013
Property, Plant and Equipment [Line Items] |  |  |
Property and equipment | $ 35,031 | $ 24,140 |
Less accumulated depreciation | (20,190) | (17,355) |
Property and equipment, net | 14,841 | 6,785 |
Depreciation expense | 3,200 | 1,800 | $ 1,200
Property, Plant and Equipment, Non-cash Additions | 1,700 |  |
Computer Equipment [Member] |  |  |
Property, Plant and Equipment [Line Items] |  |  |
Property and equipment | 3,069 | 2,596 |
Software [Member] |  |  |
Property, Plant and Equipment [Line Items] |  |  |
Property and equipment | 1,801 | 1,683 |
Furniture and Office Equipment [Member] |  |  |
Property, Plant and Equipment [Line Items] |  |  |
Property and equipment | 919 | 751 |
Lab Equipment [Member] |  |  |
Property, Plant and Equipment [Line Items] |  |  |
Property and equipment | 17,306 | 13,917 |
Leasehold Improvements [Member] |  |  |
Property, Plant and Equipment [Line Items] |  |  |
Property and equipment | $ 11,936 | $ 5,193 |

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- Definition Amount of accumulated depreciation, depletion and amortization for physical assets used in the normal conduct of business to produce goods and services. + References Reference 1: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 360 -SubTopic 10 -Section 50 -Paragraph 1 -Subparagraph (c) -URI http://asc.fasb.org/extlink&oid=6391035&loc=d3e2868-110229 Reference 2: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 210 -SubTopic 10 -Section S99 -Paragraph 1 -Subparagraph (SX 210.5-02.14) -URI http://asc.fasb.org/extlink&oid=6877327&loc=d3e13212-122682 + Details
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