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CONSOLIDATED STATEMENTS OF OPERATIONS AND COMPREHENSIVE LOSS - USD ($)
$ in Thousands
 12 Months Ended
Dec. 31, 2015
Dec. 31, 2014
Dec. 31, 2013
Revenues:      
Revenue from collaborative agreements $ 99,368 $ 47,264 $ 56,753
Revenue from government agreements 1,486 533 1,282
Total revenues 100,854 47,797 58,035
Costs and expenses:      
Research and development 98,271 70,186 46,582
General and administrative 22,765 15,926 11,087
Total costs and expenses 121,036 86,112 57,669
Income (loss) from operations (20,182) (38,315) 366
Other income (expense) 42 2 (627)
Net loss (20,140) (38,313) (261)
Other comprehensive loss:      
Unrealized loss on investments, net (5) 0 0
Comprehensive loss $ (20,145) $ (38,313) $ (261)
Basic and diluted net loss per common share (in dollars per share) $ (0.63) $ (1.40) $ (0.04)
Basic and diluted weighted average number of common shares (in shares) 31,801,645 27,384,990 6,847,697