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CONSOLIDATED STATEMENTS OF OPERATIONS AND COMPREHENSIVE INCOME (LOSS) - USD ($)
$ in Thousands
 3 Months Ended 9 Months Ended
Sep. 30, 2016
Sep. 30, 2015
Sep. 30, 2016
Sep. 30, 2015
Revenues:        
Revenue from collaborative agreements $ 2,014 $ 14,681 $ 82,404 $ 91,444
Revenue from government agreements 1,241 0 4,370 1,232
Total revenues 3,255 14,681 86,774 92,676
Costs and expenses:        
Research and development 30,296 24,103 90,982 68,227
General and administrative 7,224 6,021 20,596 16,050
Total costs and expenses 37,520 30,124 111,578 84,277
Income (loss) from operations (34,265) (15,443) (24,804) 8,399
Other income (expense) 419 1 1,059 (88)
Net income (loss) (33,846) (15,442) (23,745) 8,311
Other comprehensive income (loss):        
Unrealized gain (loss) on investments (41) 0 23 0
Comprehensive income (loss) $ (33,887) $ (15,442) $ (23,722) $ 8,311
Basic net income (loss) per common share (in dollars per share) $ (0.97) $ (0.46) $ (0.69) $ 0.27
Diluted net income (loss) per common share (in dollars per share) $ (0.97) $ (0.46) $ (0.69) $ 0.25
Basic weighted average number of common shares (in shares) 34,766,440 33,339,163 34,629,330 30,952,458
Diluted weighted average number of common shares (in shares) 34,766,440 33,339,163 34,629,330 32,960,233