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Collaboration and Other Agreements - Green Cross Corporation (Details) - USD ($)
 1 Months Ended 3 Months Ended 12 Months Ended
Jul. 31, 2016
Jun. 30, 2010
Mar. 31, 2017
Mar. 31, 2016
Dec. 31, 2014
Dec. 31, 2016
Collaboration And Other Agreements [Line Items]            
Deferred revenue included in current liabilities     $ 4,261,000     $ 4,261,000
Deferred revenue included in long-term liabilities     8,979,000     10,045,000
Green Cross            
Collaboration And Other Agreements [Line Items]            
Non-refundable upfront payment   $ 1,000,000        
Adjustment to revenue under Accounting Standards Codification         $ 1,300,000  
Proceeds from research and development services     5,500,000      
Recognized revenue under agreement     200,000 $ 100,000.0    
Deferred revenue     3,000,000     3,200,000
Deferred revenue included in current liabilities     900,000     900,000
Deferred revenue included in long-term liabilities     2,100,000     $ 2,300,000
Green Cross | Maximum            
Collaboration And Other Agreements [Line Items]            
Potential cost reimbursement for clinical trials     5,500,000.0      
Additional potential clinical and commercial milestone payments     2,500,000.0      
Janssen MGD015 Agreement | Janssen Biotech, Inc.            
Collaboration And Other Agreements [Line Items]            
Non-refundable upfront payment $ 75,000,000          
Recognized revenue under agreement     $ 100,000.0 $ 0