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CONSOLIDATED STATEMENTS OF CASH FLOWS - USD ($)
$ in Thousands
 3 Months Ended
Mar. 31, 2017
Mar. 31, 2016
Cash flows from operating activities    
Net income $ (37,655) $ (30,363)
Adjustments to reconcile net income to net cash used in operating activities:    
Depreciation and amortization expense 1,978 1,771
Share-based compensation 3,461 3,001
Changes in operating assets and liabilities:    
Accounts receivable 671 (1,287)
Prepaid expenses (1,535) 103
Other assets (81) (298)
Accounts payable (2,223) (388)
Accrued expenses 1,823 (1,788)
Lease exit liability (379) (488)
Deferred revenue (1,065) (1,893)
Deferred rent (326) (280)
Net cash used in operating activities (35,331) (31,910)
Cash flows from investing activities    
Purchases of marketable securities (56,937) (83,116)
Proceeds from sale and maturities of marketable securities 50,072 73,413
Purchases of property and equipment (1,492) (2,831)
Net cash used in investing activities (8,357) (12,534)
Cash flows from financing activities    
Proceeds from stock option exercises 108 224
Net cash provided by financing activities 108 224
Net change in cash and cash equivalents (43,580) (44,220)
Cash and cash equivalents at beginning of period 84,098 196,172
Cash and cash equivalents at end of period $ 40,518 $ 151,952