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Collaboration and Other Agreements - NIAID Contract (Details) - NIAID
 3 Months Ended 6 Months Ended
Jun. 30, 2017
USD ($)
Jun. 30, 2016
USD ($)
Jun. 30, 2017
USD ($)
Molecule
Jun. 30, 2016
USD ($)
Collaboration And Other Agreements [Line Items]        
Commercialization of molecules | Molecule     2  
Base period $ 7,500,000   $ 7,500,000  
Revenue recognized under agreement 400,000 $ 2,100,000 1,000,000 $ 3,100,000
Maximum        
Collaboration And Other Agreements [Line Items]        
Additional development funding options under agreement 17,000,000.0   17,000,000.0  
Total potential value $ 24,500,000.0   $ 24,500,000.0