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Collaboration and Other Agreements, Boehringer (Details) - Boehringer Ingelheim GmbH (Boehringer) | 1 Months Ended | 12 Months Ended
--- | --- | ---
Oct. 31, 2010 USD ($) Payment | Dec. 31, 2017 USD ($) | Dec. 31, 2016 USD ($) | Dec. 31, 2015 USD ($)
Collaboration and Other Agreements [Line Items] |  |  |  |
Non-refundable upfront payment | $ 15,000,000 |  |  |
Number of annual maintenance payments received | Payment | 3 |  |  |
Clinical milestone payments | $ 34,000,000 |  |  |
Additional potential regulatory milestone payments under agreement | 88,500,000 |  |  |
Additional potential sales milestone payments under agreement | $ 82,500,000 |  |  |
Recognized revenue under agreement |  | $ 0 | $ 0 | $ 12,500,000
Milestone payment |  |  |  | $ 5,000,000
Deferred revenue |  | $ 0 | $ 0 |

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