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Summary of Significant Accounting Policies - Adoption of 606 (Details) - USD ($) $ / shares in Units, $ in Thousands | 3 Months Ended | 12 Months Ended |
--- | --- | --- | ---
Dec. 31, 2018 | Sep. 30, 2018 | Jun. 30, 2018 | Mar. 31, 2018 | Dec. 31, 2017 | Sep. 30, 2017 | Jun. 30, 2017 | Mar. 31, 2017 | Dec. 31, 2018 | Dec. 31, 2017 | Dec. 31, 2016 | Jan. 01, 2018
Revenue, Initial Application Period Cumulative Effect Transition [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue, current | $ 21,721 |  |  |  | $ 7,202 |  |  |  | $ 21,721 | $ 7,202 |  | $ 7,742
Deferred revenue, net of current portion | 19,001 |  |  |  | 13,637 |  |  |  | 19,001 | 13,637 |  | 19,576
Accumulated deficit | (490,271) |  |  |  | (312,340) |  |  |  | (490,271) | (312,340) |  | (318,818)
Revenues | 15,794 | $ 20,798 | $ 18,834 | $ 4,695 | 152,359 | $ 1,663 | $ 1,666 | $ 2,055 | 60,121 | 157,742 | $ 91,880 |
Net loss | (44,644) | $ (34,029) | $ (43,244) | $ (49,536) | $ 105,727 | $ (47,043) | $ (40,654) | $ (37,655) | $ (171,453) | $ (19,626) | $ (58,528) |
Basic and diluted net loss per common share (in usd per share) |  |  |  |  |  |  |  |  | $ (4.19) | $ (0.54) | $ (1.69) |
Pre-Adoption |  |  |  |  |  |  |  |  |  |  |  |
Revenue, Initial Application Period Cumulative Effect Transition [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue, current | 22,210 |  |  |  |  |  |  |  | $ 22,210 |  |  | 7,202
Deferred revenue, net of current portion | 12,573 |  |  |  |  |  |  |  | 12,573 |  |  | 13,637
Accumulated deficit | (484,332) |  |  |  |  |  |  |  | (484,332) |  |  | (312,340)
Net loss |  |  |  |  |  |  |  |  | $ (171,993) |  |  |
Basic and diluted net loss per common share (in usd per share) |  |  |  |  |  |  |  |  | $ (4.20) |  |  |
Effect of Change Higher/(Lower) |  |  |  |  |  |  |  |  |  |  |  |
Revenue, Initial Application Period Cumulative Effect Transition [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue, current | (489) |  |  |  |  |  |  |  | $ (489) |  |  |
Deferred revenue, net of current portion | 6,428 |  |  |  |  |  |  |  | 6,428 |  |  |
Accumulated deficit | $ (5,939) |  |  |  |  |  |  |  | (5,939) |  |  |
Net loss |  |  |  |  |  |  |  |  | $ (540) |  |  |
Basic and diluted net loss per common share (in usd per share) |  |  |  |  |  |  |  |  | $ 0.01 |  |  |
Accounting Standards Update 2014-09 |  |  |  |  |  |  |  |  |  |  |  |
Revenue, Initial Application Period Cumulative Effect Transition [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue, current |  |  |  |  |  |  |  |  |  |  |  | 540
Deferred revenue, net of current portion |  |  |  |  |  |  |  |  |  |  |  | 5,939
Accumulated deficit |  |  |  |  |  |  |  |  |  |  |  | $ (6,478)
Revenue from collaborative agreements |  |  |  |  |  |  |  |  |  |  |  |
Revenue, Initial Application Period Cumulative Effect Transition [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  | $ 58,644 | $ 155,516 | $ 86,582 |
Revenue from collaborative agreements | Pre-Adoption |  |  |  |  |  |  |  |  |  |  |  |
Revenue, Initial Application Period Cumulative Effect Transition [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  | 58,104 |  |  |
Revenue from collaborative agreements | Effect of Change Higher/(Lower) |  |  |  |  |  |  |  |  |  |  |  |
Revenue, Initial Application Period Cumulative Effect Transition [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  | $ 540 |  |  |

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