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Collaboration and Other Agreements - Roche (Details) $ in Thousands | 1 Months Ended | 3 Months Ended |
--- | --- | --- | ---
Jan. 31, 2018 USD ($) | Dec. 31, 2017 USD ($) performance_obligation | Mar. 31, 2019 USD ($) | Mar. 31, 2018 USD ($) | Dec. 31, 2018 USD ($)
Collaboration And Other Agreements [Line Items] |  |  |  |  |
Revenues |  |  | $ 9,662 | $ 4,695 |
Deferred revenue, current |  |  | 18,085 |  | $ 21,721
Deferred revenue, net of current portion |  |  | 17,207 |  | 19,001
F. Hoffmann-La Roche Ltd. and Hoffmann-La Roche Inc. |  |  |  |  |
Collaboration And Other Agreements [Line Items] |  |  |  |  |
Non-refundable upfront payment | $ 10,000 |  |  |  |
Potential milestone payments and royalties on future sales under agreement |  | $ 370,000 |  |  |
Number of performance obligations | performance_obligation |  | 1 |  |  |
Collaborative agreement transaction price |  | $ 10,000 |  |  |
Upfront payment recognition period |  | 30 months |  |  |
Deferred revenue, current |  |  | 4,000 |  | 4,000
Deferred revenue, net of current portion |  |  | 1,000 |  | 2,000
Revenues From License Agreements | F. Hoffmann-La Roche Ltd. and Hoffmann-La Roche Inc. |  |  |  |  |
Collaboration And Other Agreements [Line Items] |  |  |  |  |
Revenues |  |  | 1,000 | $ 1,000 |
Deferred revenue |  |  | $ 5,000 |  | $ 6,000

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