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CONSOLIDATED STATEMENTS OF OPERATIONS AND COMPREHENSIVE LOSS - USD ($) $ in Thousands	3 Months Ended
	Mar. 31, 2019	Mar. 31, 2018
Revenues:
Revenues	$ 9,662	$ 4,695
Costs and expenses:
Research and development	47,060	45,670
General and administrative	10,219	9,235
Total costs and expenses	57,279	54,905
Loss from operations	(47,617)	(50,210)
Other income	2,600	674
Net loss	(45,017)	(49,536)
Other comprehensive loss:
Unrealized gain on investments	3	39
Comprehensive loss	$ (45,014)	$ (49,497)
Basic and diluted net loss per common share (in usd per share)	$ (0.99)	$ (1.34)
Basic and diluted weighted average common shares outstanding (in shares)	45,606,651	36,936,560
Revenue From Collaborative Agreements
Revenues:
Revenues	$ 9,497	$ 4,501
Revenue From Government Agreements
Revenues:
Revenues	$ 165	$ 194

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