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CONSOLIDATED STATEMENTS OF OPERATIONS AND COMPREHENSIVE LOSS - USD ($) $ in Thousands	3 Months Ended		9 Months Ended
	Sep. 30, 2019	Sep. 30, 2018	Sep. 30, 2019	Sep. 30, 2018
Revenues:
Revenues	$ 18,741	$ 20,798	$ 38,996	$ 44,327
Costs and expenses:
Research and development	44,852	46,218	143,352	143,902
General and administrative	11,833	9,584	34,174	29,953
Total costs and expenses	56,685	55,802	177,526	173,855
Loss from operations	(37,944)	(35,004)	(138,530)	(129,528)
Other income (expense)	(6,687)	975	17,115	2,719
Net loss	(44,631)	(34,029)	(121,415)	(126,809)
Other comprehensive income:
Unrealized gain (loss) on investments	(11)	(18)	26	61
Comprehensive loss	$ (44,642)	$ (34,047)	$ (121,389)	$ (126,748)
Basic and diluted net loss per common share (in usd per share)	$ (0.91)	$ (0.81)	$ (2.54)	$ (3.13)
Basic and diluted weighted average common shares outstanding (in shares)	48,902,766	42,239,327	47,796,957	40,462,658
Revenue From Collaborative Agreements
Revenues:
Revenues	$ 17,984	$ 20,617	$ 37,468	$ 43,670
Revenue From Government Agreements
Revenues:
Revenues	$ 757	$ 181	$ 1,528	$ 657

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