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Collaboration and Other Agreements, NIAID Contract (Details) |  | 3 Months Ended | 12 Months Ended
--- | --- | --- | ---
Sep. 15, 2015 USD ($) Molecule | Dec. 31, 2019 USD ($) | Sep. 30, 2019 USD ($) | Jun. 30, 2019 USD ($) | Mar. 31, 2019 USD ($) | Dec. 31, 2018 USD ($) | Sep. 30, 2018 USD ($) | Jun. 30, 2018 USD ($) | Mar. 31, 2018 USD ($) | Dec. 31, 2019 USD ($) | Dec. 31, 2018 USD ($) | Dec. 31, 2017 USD ($)
Collaboration and Other Agreements [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  | $ 25,192,000 | $ 18,741,000 | $ 10,593,000 | $ 9,662,000 | $ 15,794,000 | $ 20,798,000 | $ 18,834,000 | $ 4,695,000 | $ 64,188,000 | $ 60,121,000 | $ 157,742,000
NIAID |  |  |  |  |  |  |  |  |  |  |  |
Collaboration and Other Agreements [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Commercialization of molecules | Molecule | 2 |  |  |  |  |  |  |  |  |  |  |
Base period value | $ 7,500,000 |  |  |  |  |  |  |  |  |  |  |
Proceeds from additional development funding options under agreement |  |  |  |  |  |  |  |  |  |  |  | 10,800,000
NIAID | Maximum |  |  |  |  |  |  |  |  |  |  |  |
Collaboration and Other Agreements [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Additional development funding options | 17,000,000.0 |  |  |  |  |  |  |  |  |  |  |
Total potential value | $ 24,500,000 |  |  |  |  |  |  |  |  |  |  |
Revenues From License Agreements | NIAID |  |  |  |  |  |  |  |  |  |  |  |
Collaboration and Other Agreements [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  | $ 2,200,000 | $ 1,300,000 | $ 1,700,000

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