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Collaboration and Other Agreements, Roche (Details) | 1 Months Ended | 3 Months Ended | 12 Months Ended
--- | --- | --- | ---
Jan. 31, 2018 USD ($) | Dec. 31, 2017 USD ($) performance_obligation | Dec. 31, 2019 USD ($) | Sep. 30, 2019 USD ($) | Jun. 30, 2019 USD ($) | Mar. 31, 2019 USD ($) | Dec. 31, 2018 USD ($) | Sep. 30, 2018 USD ($) | Jun. 30, 2018 USD ($) | Mar. 31, 2018 USD ($) | Dec. 31, 2019 USD ($) | Dec. 31, 2018 USD ($) | Dec. 31, 2017 USD ($) performance_obligation
Collaboration and Other Agreements [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  | $ 25,192,000 | $ 18,741,000 | $ 10,593,000 | $ 9,662,000 | $ 15,794,000 | $ 20,798,000 | $ 18,834,000 | $ 4,695,000 | $ 64,188,000 | $ 60,121,000 | $ 157,742,000
Deferred revenue |  | $ 27,318,000 |  |  |  |  | 40,722,000 |  |  |  |  | 40,722,000 | $ 27,318,000
Deferred revenue, current |  |  | 10,700,000 |  |  |  | 21,721,000 |  |  |  | 10,700,000 | 21,721,000 |
Deferred revenue included in long-term liabilities |  |  | 9,153,000 |  |  |  | 19,001,000 |  |  |  | 9,153,000 | 19,001,000 |
Roche |  |  |  |  |  |  |  |  |  |  |  |  |
Collaboration and Other Agreements [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Non-refundable upfront payment | $ 10,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |
Potential milestone payments and royalties on future sales |  |  | 370,000,000.0 |  |  |  |  |  |  |  | 370,000,000.0 |  |
Number of performance obligations | performance_obligation |  | 1 |  |  |  |  |  |  |  |  |  |  | 1
Transaction price |  | $ 10,000,000.0 |  |  |  |  |  |  |  |  |  |  | $ 10,000,000.0
Expected period of development |  | 30 months |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  | 6,000,000.0 | 4,000,000.0 |
Deferred revenue |  |  | $ 0 |  |  |  |  |  |  |  | $ 0 |  |
Deferred revenue |  |  |  |  |  |  | 6,000,000.0 |  |  |  |  | 6,000,000.0 |
Deferred revenue, current |  |  |  |  |  |  | 4,000,000.0 |  |  |  |  | 4,000,000.0 |
Deferred revenue included in long-term liabilities |  |  |  |  |  |  | $ 2,000,000.0 |  |  |  |  | $ 2,000,000.0 |

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