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CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-In Capital | Accumulated Deficit | Accumulated Other Comprehensive Income
--- | --- | --- | --- | --- | ---
Beginning balance (in shares) at Dec. 31, 2018 |  | 42,353,301 |  |  |
Beginning balance at Dec. 31, 2018 | $ 242,877 | $ 424 | $ 732,727 | $ (490,271) | $ (3)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Share-based compensation | 3,750 |  | 3,750 |  |
Issuance of common stock, net of offering costs (in shares) |  | 6,325,000 |  |  |
Issuance of common stock, net of offering costs | 118,657 | $ 63 | 118,594 |  |
Stock plan related activity (in shares) |  | 126,707 |  |  |
Stock plan related activity | 347 | $ 1 | 346 |  |
Unrealized gain (loss) on investments | 3 |  |  |  | 3
Net loss | (45,017) |  |  | (45,017) |
Ending balance (in shares) at Mar. 31, 2019 |  | 48,805,008 |  |  |
Ending balance at Mar. 31, 2019 | 320,617 | $ 488 | 855,417 | (535,288) | 0
Beginning balance (in shares) at Dec. 31, 2019 |  | 48,958,763 |  |  |
Beginning balance at Dec. 31, 2019 | 230,628 | $ 490 | 872,204 | (642,082) | 16
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Share-based compensation | $ 4,451 |  | 4,451 |  |
Stock plan related activity (in shares) | 1,403,627 | 172,387 |  |  |
Stock plan related activity | $ 162 | $ 2 | 160 |  |
Unrealized gain (loss) on investments | 56 |  |  |  | 56
Net loss | (44,724) |  |  | (44,724) |
Ending balance (in shares) at Mar. 31, 2020 |  | 49,131,150 |  |  |
Ending balance at Mar. 31, 2020 | $ 190,573 | $ 492 | $ 876,815 | $ (686,806) | $ 72

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