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CONSOLIDATED STATEMENTS OF OPERATIONS AND COMPREHENSIVE LOSS - USD ($) $ in Thousands	3 Months Ended
	Mar. 31, 2020	Mar. 31, 2019
Revenues:
Revenues	$ 13,682	$ 9,662
Costs and expenses:
Research and development	48,894	47,060
General and administrative	10,233	10,219
Total costs and expenses	59,127	57,279
Loss from operations	(45,445)	(47,617)
Other income	721	2,600
Net loss	(44,724)	(45,017)
Other comprehensive income:
Unrealized gain on investments	56	3
Comprehensive loss	$ (44,668)	$ (45,014)
Basic and diluted net loss per common share (in usd per share)	$ (0.91)	$ (0.99)
Basic and diluted weighted average common shares outstanding (in shares)	49,012,663	45,606,651
Revenue From Collaborative Agreements
Revenues:
Revenues	$ 12,967	$ 9,497
Revenue From Government Agreements
Revenues:
Revenues	$ 715	$ 165

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