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CONSOLIDATED STATEMENTS OF OPERATIONS AND COMPREHENSIVE LOSS - USD ($)
$ in Thousands
 3 Months Ended 6 Months Ended
Jun. 30, 2020
Jun. 30, 2019
Jun. 30, 2020
Jun. 30, 2019
Revenues:        
Revenues $ 20,257 $ 10,593 $ 33,939 $ 20,255
Costs and expenses:        
Research and development 57,351 51,440 106,245 98,500
General and administrative 10,216 12,122 20,449 22,341
Total costs and expenses 67,567 63,562 126,694 120,841
Loss from operations (47,310) (52,969) (92,755) (100,586)
Other income 425 21,202 1,146 23,802
Net loss (46,885) (31,767) (91,609) (76,784)
Other comprehensive loss:        
Unrealized gain (loss) on investments (55) 34 1 37
Comprehensive loss $ (46,940) $ (31,733) $ (91,608) $ (76,747)
Basic and diluted net loss per common share (in usd per share) $ (0.94) $ (0.65) $ (1.85) $ (1.63)
Basic and diluted weighted average common shares outstanding (in shares) 50,018,462 48,845,234 49,515,562 47,234,889
Revenue From Collaborative Agreements        
Revenues:        
Revenues $ 15,636 $ 9,987 $ 28,603 $ 19,484
Revenue From Government Agreements        
Revenues:        
Revenues $ 4,621 $ 606 $ 5,336 $ 771