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Collaboration and Other Agreements - NIAID Contract (Details)
 3 Months Ended 9 Months Ended 12 Months Ended
Sep. 15, 2015
USD ($)
Molecule
Sep. 30, 2020
USD ($)
Sep. 30, 2019
USD ($)
Sep. 30, 2020
USD ($)
Sep. 30, 2019
USD ($)
Dec. 31, 2017
USD ($)
Collaboration And Other Agreements [Line Items]            
Revenues   $ 18,253,000 $ 18,741,000 $ 52,192,000 $ 38,996,000  
NIAID            
Collaboration And Other Agreements [Line Items]            
Commercialization of molecules | Molecule 2          
Base period $ 7,500,000          
Additional development funding options under agreement 17,000,000.0          
Total potential value $ 24,500,000          
Proceeds from additional development funding options under agreement           $ 10,800,000
Revenues From Grants | NIAID            
Collaboration And Other Agreements [Line Items]            
Revenues   $ 800,000 $ 800,000 $ 6,200,000 $ 1,500,000