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CONSOLIDATED STATEMENTS OF OPERATIONS AND COMPREHENSIVE LOSS - USD ($) $ in Thousands	3 Months Ended		9 Months Ended
	Sep. 30, 2020	Sep. 30, 2019	Sep. 30, 2020	Sep. 30, 2019
Revenues:
Revenues	$ 18,253	$ 18,741	$ 52,192	$ 38,996
Costs and expenses:
Research and development	44,656	44,852	150,901	143,352
General and administrative	9,732	11,833	30,181	34,174
Total costs and expenses	54,388	56,685	181,082	177,526
Loss from operations	(36,135)	(37,944)	(128,890)	(138,530)
Other income (expense)	92	(6,687)	1,238	17,115
Net loss	(36,043)	(44,631)	(127,652)	(121,415)
Other comprehensive loss:
Unrealized gain (loss) on investments	(15)	(11)	(14)	26
Comprehensive loss	$ (36,058)	$ (44,642)	$ (127,666)	$ (121,389)
Basic and diluted net loss per common share (in usd per share)	$ (0.66)	$ (0.91)	$ (2.49)	$ (2.54)
Basic and diluted weighted average common shares outstanding (in shares)	54,463,412	48,902,766	51,176,884	47,796,957
Revenue From Collaborative Agreements
Revenues:
Revenues	$ 17,415	$ 17,984	$ 46,018	$ 37,468
Revenue From Government Agreements
Revenues:
Revenues	$ 838	$ 757	$ 6,174	$ 1,528

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