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CONSOLIDATED STATEMENTS OF CASH FLOWS - USD ($)
$ in Thousands
 9 Months Ended
Sep. 30, 2020
Sep. 30, 2019
Cash flows from operating activities    
Net loss $ (127,652) $ (121,415)
Adjustments to reconcile net loss to net cash used in operating activities:    
Depreciation expense 9,067 9,238
Amortization of premiums and discounts on marketable securities (387) (1,119)
Stock-based compensation 15,419 14,081
Changes in operating assets and liabilities:    
Accounts receivable (6,033) 20,080
Prepaid expenses and other current assets 654 (5,612)
Other assets 1,615 (2,621)
Accounts payable 137 (913)
Accrued expenses 5,499 (4,678)
Lease liabilities (2,113) 3,847
Deferred revenue (4,816) (7,114)
Other liabilities 2,245 0
Net cash used in operating activities (106,365) (96,226)
Cash flows from investing activities    
Purchases of marketable securities (151,230) (214,178)
Proceeds from sale and maturities of marketable securities 161,866 130,236
Purchases of property and equipment (2,346) (3,042)
Net cash provided by (used in) investing activities 8,290 (86,984)
Cash flows from financing activities    
Proceeds from issuance of common stock, net of offering costs 170,494 118,657
Proceeds from stock option exercises and ESPP purchases 4,757 1,018
Retirement Of Treasury Stock (2,011) 0
Net cash provided by financing activities 173,240 119,675
Net change in cash and cash equivalents 75,165 (63,535)
Cash and cash equivalents at beginning of period 126,472 220,128
Cash and cash equivalents at end of period 201,637 156,593
Supplemental Cash Flow Information [Abstract]    
Right-of-use assets modified in exchange for operating lease obligations $ 0 $ 6,408