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Collaboration and Other Agreements - Roche (Details) | 1 Months Ended | 12 Months Ended
--- | --- | ---
Jan. 31, 2018 USD ($) | Dec. 31, 2017 | Dec. 31, 2020 USD ($) | Dec. 31, 2019 USD ($) | Dec. 31, 2018 USD ($) performanceObligation
Collaboration and Other Agreements [Line Items] |  |  |  |  |
Revenues |  |  | $ 104,883,000 | $ 64,188,000 | $ 60,121,000
Deferred revenue, current |  |  | 4,456,000 | 10,700,000 |
Deferred revenue included in long-term liabilities |  |  | 6,926,000 | 9,153,000 |
Roche |  |  |  |  |
Collaboration and Other Agreements [Line Items] |  |  |  |  |
Non-refundable upfront payment | $ 10,000,000.0 |  |  |  |
Potential milestone payments and royalties on future sales |  |  | 370,000,000.0 |  |
Number of performance obligations | performanceObligation |  |  |  |  | 1
Transaction price |  |  |  |  | $ 10,000,000.0
Expected period of development |  | 30 months |  |  |
Revenues |  |  | 6,000,000.0 | $ 4,000,000.0 |
Deferred revenue |  |  | $ 0 |  |

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