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CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-In Capital | Accumulated Deficit | Accumulated Other Comprehensive Income (Loss)
--- | --- | --- | --- | --- | ---
Beginning balance (in shares) at Dec. 31, 2019 |  | 48,958,763 |  |  |
Beginning balance at Dec. 31, 2019 | $ 230,628 | $ 490 | $ 872,204 | $ (642,082) | $ 16
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Share-based compensation | 4,451 |  | 4,451 |  |
Stock plan related activity (in shares) |  | 172,387 |  |  |
Stock plan related activity | 162 | $ 2 | 160 |  |
Unrealized gain (loss) on investments | 56 |  |  |  | 56
Net loss | (44,724) |  |  | (44,724) |
Ending balance (in shares) at Mar. 31, 2020 |  | 49,131,150 |  |  |
Ending balance at Mar. 31, 2020 | 190,573 | $ 492 | 876,815 | (686,806) | 72
Beginning balance (in shares) at Dec. 31, 2019 |  | 48,958,763 |  |  |
Beginning balance at Dec. 31, 2019 | 230,628 | $ 490 | 872,204 | (642,082) | 16
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Stock plan related activity |  | $ 2 |  |  |
Net loss | (91,609) |  |  |  |
Ending balance (in shares) at Jun. 30, 2020 |  | 53,365,003 |  |  |
Ending balance at Jun. 30, 2020 | 247,784 | $ 534 | 980,924 | (733,691) | 17
Beginning balance (in shares) at Mar. 31, 2020 |  | 49,131,150 |  |  |
Beginning balance at Mar. 31, 2020 | 190,573 | $ 492 | 876,815 | (686,806) | 72
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Share-based compensation | 5,136 |  | 5,136 |  |
Net Income (Loss), Including Portion Attributable to Noncontrolling Interest | (46,885) |  |  | (46,885) |
Issuance of common stock, net of offering costs (in shares) |  | 4,060,482 |  |  |
Issuance of common stock, net of offering costs | 96,512 | $ 40 | 96,472 |  |
Stock plan related activity (in shares) |  | 173,371 |  |  |
Stock plan related activity | 2,503 |  | 2,501 |  |
Unrealized gain (loss) on investments | (55) |  |  |  | (55)
Net loss | (46,885) |  |  |  |
Ending balance (in shares) at Jun. 30, 2020 |  | 53,365,003 |  |  |
Ending balance at Jun. 30, 2020 | 247,784 | $ 534 | 980,924 | (733,691) | 17
Beginning balance (in shares) at Dec. 31, 2020 |  | 56,244,771 |  |  |
Beginning balance at Dec. 31, 2020 | 295,884 | $ 562 | 1,067,150 | (771,821) | (7)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Share-based compensation | 5,243 |  | 5,243 |  |
Issuance of common stock, net of offering costs (in shares) |  | 3,622,186 |  |  |
Issuance of common stock, net of offering costs | 98,200 | $ 36 | 98,164 |  |
Stock plan related activity (in shares) |  | 144,249 |  |  |
Stock plan related activity | 2,458 | $ 2 | 2,456 |  |
Unrealized gain (loss) on investments | 18 |  |  |  | 18
Net loss | (51,272) |  |  | (51,272) |
Ending balance (in shares) at Mar. 31, 2021 |  | 60,011,206 |  |  |
Ending balance at Mar. 31, 2021 | 350,531 | $ 600 | 1,173,013 | (823,093) | 11
Beginning balance (in shares) at Dec. 31, 2020 |  | 56,244,771 |  |  |
Beginning balance at Dec. 31, 2020 | $ 295,884 | $ 562 | 1,067,150 | (771,821) | (7)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Stock plan related activity (in shares) | 1,790,246 |  |  |  |
Net loss | $ (91,207) |  |  |  |
Ending balance (in shares) at Jun. 30, 2021 |  | 60,133,447 |  |  |
Ending balance at Jun. 30, 2021 | 319,045 | $ 601 | 1,181,471 | (863,028) | 1
Beginning balance (in shares) at Mar. 31, 2021 |  | 60,011,206 |  |  |
Beginning balance at Mar. 31, 2021 | 350,531 | $ 600 | 1,173,013 | (823,093) | 11
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Share-based compensation | 6,113 |  | 6,113 |  |
Net Income (Loss), Including Portion Attributable to Noncontrolling Interest | (39,935) |  |  | (39,935) |
Stock plan related activity (in shares) |  | 122,241 |  |  |
Stock plan related activity | 2,346 | $ 1 | 2,345 |  |
Unrealized gain (loss) on investments | (10) |  |  |  | (10)
Net loss | (39,935) |  |  |  |
Ending balance (in shares) at Jun. 30, 2021 |  | 60,133,447 |  |  |
Ending balance at Jun. 30, 2021 | $ 319,045 | $ 601 | $ 1,181,471 | $ (863,028) | $ 1

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