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Collaboration and Other Agreements - NIAID Contract (Details) $ in Thousands		3 Months Ended		9 Months Ended
	Sep. 15, 2015 Molecule	Sep. 30, 2021 USD ($)	Sep. 30, 2020 USD ($)	Sep. 30, 2021 USD ($)	Sep. 30, 2020 USD ($)
Collaboration And Other Agreements [Line Items]
Revenues		$ 15,662	$ 18,253	$ 63,300	$ 52,192
NIAID
Collaboration And Other Agreements [Line Items]
Number of commercialized molecules | Molecule	2
Total potential value		25,100
Revenues From Grants | NIAID
Collaboration And Other Agreements [Line Items]
Revenues		$ 100	$ 800	$ 1,300	$ 6,200

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