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Stock-Based Compensation - Stock-Based Compensation Expense (Details) - USD ($) $ / shares in Units, $ in Thousands | 1 Months Ended | 3 Months Ended |  |
--- | --- | --- | --- | ---
May 31, 2017 | Mar. 31, 2022 | Mar. 31, 2021 | Dec. 31, 2021 | Oct. 31, 2013
Share-based Compensation Arrangement by Share-based Payment Award [Line Items] |  |  |  |  |
Offering period | 6 months |  |  |  |
Options outstanding (in shares) |  | 10,506,580 |  | 8,373,921 |
Weighted-average exercise price of stock options outstanding (in dollars per share) |  | $ 19.08 |  | $ 21.47 |
Employee Service Share-based Compensation, Allocation of Recognized Period Costs [Line Items] |  |  |  |  |
Stock-based compensation expense |  | $ 5,274 | $ 5,286 |  |
2016 Employee Stock Purchase Plan |  |  |  |  |
Share-based Compensation Arrangement by Share-based Payment Award [Line Items] |  |  |  |  |
Share-based compensation, number of shares authorized (in shares) | 800,000 |  |  |  |
Payroll deduction percentage | 10.00% |  |  |  |
Purchase price of common stock percent of the fair market value | 85.00% |  |  |  |
Common stock purchased (in shares) |  | 0 |  |  |
2003 Stock Incentive Plan |  |  |  |  |
Share-based Compensation Arrangement by Share-based Payment Award [Line Items] |  |  |  |  |
Options outstanding (in shares) |  | 160,357 |  |  |
Stock Incentive Plan 2013 |  |  |  |  |
Share-based Compensation Arrangement by Share-based Payment Award [Line Items] |  |  |  |  |
Share-based compensation, number of shares authorized (in shares) |  | 15,816,949 |  |  |
Options outstanding (in shares) |  | 10,346,223 |  |  |
Potential annual increase in shares reserved for future issuance (in shares) |  |  |  |  | 1,960,168
Potential annual increase in shares reserved for future issuance as percentage of outstanding share |  | 4.00% |  |  |
Research and Development |  |  |  |  |
Employee Service Share-based Compensation, Allocation of Recognized Period Costs [Line Items] |  |  |  |  |
Stock-based compensation expense |  | $ 2,392 | 2,727 |  |
General and Administrative |  |  |  |  |
Employee Service Share-based Compensation, Allocation of Recognized Period Costs [Line Items] |  |  |  |  |
Stock-based compensation expense |  | $ 2,882 | $ 2,559 |  |

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