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Collaboration and Other Agreements - NIAID Contract (Details) $ in Thousands		3 Months Ended
	Sep. 15, 2015 Molecule	Mar. 31, 2022 USD ($)	Mar. 31, 2021 USD ($)
Collaboration And Other Agreements [Line Items]
Revenues		$ 11,101	$ 16,881
NIAID
Collaboration And Other Agreements [Line Items]
Number of commercialized molecules | Molecule	2
Total potential value		25,100
Revenues From Grants | NIAID
Collaboration And Other Agreements [Line Items]
Revenues		$ 400	$ 800

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