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Stock-Based Compensation - Stock Option Activity (Details) - USD ($)
$ / shares in Units, $ in Thousands
3 Months Ended 6 Months Ended
Mar. 31, 2022
Mar. 31, 2021
Jun. 30, 2021
Shares      
Shares, Outstanding, Beginning Balance (in shares) 8,373,921    
Shares, Granted (in shares) 2,349,197    
Shares, Exercised (in shares) (25,646)    
Shares, Forfeited or expired (in shares) (157,197)    
Share-based Compensation Arrangement by Share-based Payment Award, Options, Expirations in Period (33,695)    
Shares, Outstanding, Ending Balance (in shares) 10,506,580    
Shares, Exercisable (in shares) 5,853,827    
Shares, Vested and expected to vest (in shares) 9,693,151    
Weighted- Average Exercise Price      
Weighted- Average Exercise Price, Outstanding, Beginning Balance (in dollars per share) $ 21.47    
Weighted- Average Exercise Price, Granted (in dollars per share) 10.21    
Weighted- Average Exercise Price, Exercised (in dollars per share) 1.44    
Weighted- Average Exercise Price, Forfeited (in dollars per share) 16.51    
Weighted- Average Exercise Price, Expired (in dollars per share) 21.16    
Weighted- Average Exercise Price, Outstanding, Ending Balance (in dollars per share) 19.08    
Weighted- Average Exercise Price, Exercisable (in dollars per share) 22.61    
Weighted- Average Exercise Price, Vested and expected to vest (in dollars per share) $ 19.41    
Share-based Compensation Arrangement by Share-based Payment Award, Options, Additional Disclosures [Abstract]      
Weighted- Average Remaining Contractual Term, Outstanding (in years) 7 years 1 month 6 days   6 years 7 months 6 days
Weighted- Average Remaining Contractual Term, Exercisable (in years) 5 years 6 months    
Weighted- Average Remaining Contractual Term, Vested and expected to vest (in years) 7 years    
Aggregate Intrinsic Value, Outstanding, Ending Balance $ 904    
Aggregate Intrinsic Value, Exercisable 898    
Aggregate Intrinsic Value, Vested and expected to vest $ 903    
Weighted-average grant-date fair value of options granted (in dollars per share) $ 7.48 $ 14.23  
Intrinsic value of options exercised $ 300 $ 1,400  
Cash received for options exercised   2,500  
Fair value of shares vested 5,400 $ 4,100  
Unrecognized compensation expense related to non-vested stock-options, net of related forfeiture estimates $ 38,400    
Unrecognized compensation expense recognition period (in years) 1 year 7 months 6 days