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CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY - USD ($)
$ in Thousands
Total
Common Stock
Additional Paid-In Capital
Accumulated Deficit
Accumulated Other Comprehensive Income (Loss)
Beginning balance (in shares) at Dec. 31, 2020   56,244,771      
Beginning balance at Dec. 31, 2020 $ 295,884 $ 562 $ 1,067,150 $ (771,821) $ (7)
Increase (Decrease) in Stockholders' Equity [Roll Forward]          
Share-based compensation 5,243   5,243    
Stock plan activity (in shares)   144,249      
Stock plan related activity 2,458 $ 2 2,456    
Issuance of common stock, net of offering costs (in shares)   3,622,186      
Issuance of common stock, net of offering costs 98,200 $ 36 98,164    
Unrealized gain (loss) on investments 18       18
Net loss (51,272)     (51,272)  
Ending balance (in shares) at Mar. 31, 2021   60,011,206      
Ending balance at Mar. 31, 2021 350,531 $ 600 1,173,013 (823,093) 11
Beginning balance (in shares) at Dec. 31, 2021   61,307,428      
Beginning balance at Dec. 31, 2021 239,618 $ 613 1,213,002 (973,936) (61)
Increase (Decrease) in Stockholders' Equity [Roll Forward]          
Share-based compensation 5,224   5,224    
Stock plan activity (in shares)   25,646      
Stock plan related activity 37   37    
Unrealized gain (loss) on investments (222)       (222)
Net loss (66,443)     (66,443)  
Ending balance (in shares) at Mar. 31, 2022   61,333,074      
Ending balance at Mar. 31, 2022 $ 178,214 $ 613 $ 1,218,263 $ (1,040,379) $ (283)