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Income Taxes - Components of the Company's Deferred Tax Assets (Liabilities) (Details) - USD ($) $ in Thousands | Dec. 31, 2022 | Dec. 31, 2021
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Deferred income tax assets: |  |
Federal U.S. net operating loss carryforward | $ 163,071 | $ 175,802
State net operating loss carryforward | 44,784 | 49,965
Research and development credit, net | 65,084 | 60,514
Orphan drug credit, net | 35,703 | 24,858
Operating lease liabilities | 9,585 | 7,008
Deferred revenue | 0 | 1,245
Section 174 deferred tax asset | 43,192 | 0
Depreciation | 158 | 0
Other | 19,733 | 16,727
Gross deferred income tax assets | 381,310 | 336,119
Valuation allowance | (372,267) | (327,595)
Net deferred income tax assets | 9,043 | 8,524
Deferred income tax liabilities: |  |
Depreciation | 0 | (1,688)
Operating lease ROU assets | (7,522) | (4,576)
Prepaid expenditures | (1,521) | (2,260)
Gross deferred income tax liabilities | (9,043) | (8,524)
Net deferred income tax asset/(liability) | $ 0 | $ 0

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