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Revenue - NIAID Contract (Details)	12 Months Ended
	Dec. 31, 2022 USD ($)	Dec. 31, 2021 USD ($)	Dec. 31, 2020 USD ($)	Oct. 31, 2015 productCandidate
Collaboration and Other Agreements [Line Items]
Revenues	$ 151,941,000	$ 77,447,000	$ 104,883,000
NIAID
Collaboration and Other Agreements [Line Items]
Total potential value	25,100,000
Boehringer Ingelheim International GmbH | Boehringer Collaboration and License Agreement
Collaboration and Other Agreements [Line Items]
Number of product candidates to develop | productCandidate				2
RevenuesFromGrantsMember | NIAID
Collaboration and Other Agreements [Line Items]
Number of product candidates to develop		2
Revenues	$ 1,900,000	$ 1,800,000	$ 7,100,000

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