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CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-In Capital | Accumulated Deficit | Accumulated Other Comprehensive Income (Loss)
--- | --- | --- | --- | --- | ---
Beginning balance (in shares) at Dec. 31, 2021 |  | 61,307,428 |  |  |
Beginning balance at Dec. 31, 2021 | $ 239,618 | $ 613 | $ 1,213,002 | $ (973,936) | $ (61)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Share-based compensation | 5,224 |  | 5,224 |  |
Stock plan activity (in shares) |  | 25,646 |  |  |
Stock plan related activity | 37 |  | 37 |  |
Unrealized gain (loss) on investments | (222) |  |  |  | (222)
Net loss | (66,443) |  |  | (66,443) |
Ending balance (in shares) at Mar. 31, 2022 |  | 61,333,074 |  |  |
Ending balance at Mar. 31, 2022 | 178,214 | $ 613 | 1,218,263 | (1,040,379) | (283)
Beginning balance (in shares) at Dec. 31, 2021 |  | 61,307,428 |  |  |
Beginning balance at Dec. 31, 2021 | 239,618 | $ 613 | 1,213,002 | (973,936) | (61)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Net loss | (132,560) |  |  |  |
Ending balance (in shares) at Sep. 30, 2022 |  | 61,462,189 |  |  |
Ending balance at Sep. 30, 2022 | 122,958 | $ 615 | 1,228,952 | (1,106,496) | (113)
Beginning balance (in shares) at Mar. 31, 2022 |  | 61,333,074 |  |  |
Beginning balance at Mar. 31, 2022 | 178,214 | $ 613 | 1,218,263 | (1,040,379) | (283)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Share-based compensation | 5,350 |  | 5,350 |  |
Stock plan related activity | 264 | $ 2 | 262 |  |
Shares, Granted (in shares) |  | 125,716 |  |  |
Unrealized gain (loss) on investments | (43) |  |  |  | (43)
Net loss | (41,304) |  |  | (41,304) |
Ending balance (in shares) at Jun. 30, 2022 |  | 61,458,790 |  |  |
Ending balance at Jun. 30, 2022 | 142,481 | $ 615 | 1,223,875 | (1,081,683) | (326)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Share-based compensation | 5,077 |  | 5,077 |  |
Shares, Granted (in shares) |  | 3,399 |  |  |
Unrealized gain (loss) on investments | 213 |  |  |  | 213
Net loss | (24,813) |  |  | (24,813) |
Ending balance (in shares) at Sep. 30, 2022 |  | 61,462,189 |  |  |
Ending balance at Sep. 30, 2022 | 122,958 | $ 615 | 1,228,952 | (1,106,496) | (113)
Beginning balance (in shares) at Dec. 31, 2022 |  | 61,701,467 |  |  |
Beginning balance at Dec. 31, 2022 | 142,013 | $ 617 | 1,235,095 | (1,093,694) | (5)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Share-based compensation | 4,788 |  | 4,788 |  |
Stock plan activity (in shares) |  | 42,098 |  |  |
Stock plan related activity | (154) |  | (154) |  |
Issuance of common stock, net of offering costs (in shares) |  | 95,000 |  |  |
Issuance of common stock, net of offering costs | 617 | $ 1 | 616 |  |
Unrealized gain (loss) on investments | 13 |  |  |  | 13
Net loss | (38,009) |  |  | (38,009) |
Ending balance (in shares) at Mar. 31, 2023 |  | 61,838,565 |  |  |
Ending balance at Mar. 31, 2023 | 109,268 | $ 618 | 1,240,345 | (1,131,703) | 8
Beginning balance (in shares) at Dec. 31, 2022 |  | 61,701,467 |  |  |
Beginning balance at Dec. 31, 2022 | $ 142,013 | $ 617 | 1,235,095 | (1,093,694) | (5)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Shares, Granted (in shares) | 3,267,727 |  |  |  |
Net loss | $ 37,014 |  |  |  |
Ending balance (in shares) at Sep. 30, 2023 |  | 62,028,904 |  |  |
Ending balance at Sep. 30, 2023 | 193,980 | $ 620 | 1,250,074 | (1,056,680) | (34)
Beginning balance (in shares) at Mar. 31, 2023 |  | 61,838,565 |  |  |
Beginning balance at Mar. 31, 2023 | 109,268 | $ 618 | 1,240,345 | (1,131,703) | 8
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Share-based compensation | 4,436 |  | 4,436 |  |
Stock plan related activity | 216 | $ 1 | 215 |  |
Shares, Granted (in shares) |  | 63,793 |  |  |
Issuance of common stock, net of offering costs (in shares) |  | 36,135 |  |  |
Issuance of common stock, net of offering costs | 235 |  | 235 |  |
Unrealized gain (loss) on investments | (80) |  |  |  | (80)
Net loss | 57,469 |  |  | 57,469 |
Ending balance (in shares) at Jun. 30, 2023 |  | 61,938,493 |  |  |
Ending balance at Jun. 30, 2023 | 171,544 | $ 619 | 1,245,231 | (1,074,234) | (72)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Share-based compensation | 4,746 |  | 4,746 |  |
Stock plan related activity | (89) | $ 1 |  |  |
Shares, Granted (in shares) |  | 54,276 |  |  |
Stock plan related activity |  |  | (90) |  |
Issuance of common stock, net of offering costs (in shares) |  | 36,135 |  |  |
Issuance of common stock, net of offering costs | 187 |  | 187 |  |
Unrealized gain (loss) on investments | 38 |  |  |  | 38
Net loss | 17,554 |  |  | 17,554 |
Ending balance (in shares) at Sep. 30, 2023 |  | 62,028,904 |  |  |
Ending balance at Sep. 30, 2023 | $ 193,980 | $ 620 | $ 1,250,074 | $ (1,056,680) | $ (34)

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