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Income Taxes - Components of the Company's Deferred Tax Assets (Liabilities) (Details) - USD ($) $ in Thousands | Dec. 31, 2024 | Dec. 31, 2023 | Dec. 31, 2022 | Dec. 31, 2021
--- | --- | --- | --- | ---
Deferred income tax assets: |  |  |  |
Federal U.S. net operating loss carryforward | $ 116,341 | $ 140,706 |  |
State net operating loss carryforward | 31,758 | 33,350 |  |
Research and development credit, net | 79,108 | 68,251 |  |
Orphan drug credit, net | 33,834 | 33,330 |  |
Operating lease liabilities | 10,308 | 9,078 |  |
Deferred revenue | 18,795 | 17,442 |  |
Section 174 deferred tax asset | 98,810 | 48,421 |  |
Depreciation | 16,272 | 16,217 |  |
Other | 4,971 | 6,617 |  |
Gross deferred income tax assets | 410,197 | 373,412 |  |
Valuation allowance | (401,297) | (365,010) | $ (372,267) | $ (327,592)
Net deferred income tax assets | 8,900 | 8,402 |  |
Deferred income tax liabilities: |  |  |  |
Operating lease ROU assets | (6,745) | (6,372) |  |
Prepaid expenditures | (2,155) | (2,030) |  |
Gross deferred income tax liabilities | (8,900) | (8,402) |  |
Net deferred income tax asset/(liability) | $ 0 | $ 0 |  |

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