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License Agreements - Additional Information (Details) - USD ($)
$ in Thousands
1 Months Ended 3 Months Ended 9 Months Ended
Jun. 30, 2018
Sep. 30, 2016
Mar. 31, 2016
Sep. 30, 2023
Sep. 30, 2022
Sep. 30, 2023
Sep. 30, 2022
License Agreements [Line Items]              
Research and development expense       $ 6,112 $ 8,624 $ 21,135 $ 27,576
Stanford License Agreements [Member]              
License Agreements [Line Items]              
Research and development expense           100 100
Stanford License Agreements [Member] | Stanford [Member]              
License Agreements [Line Items]              
Payments for milestone agreement or earned royalties on achievement of milestones     $ 5,000        
Payments for achievement of specified development and regulatory milestones $ 400            
Payments for milestone agreement on achievement of milestones           0  
Stanford License Agreements [Member] | Stanford [Member] | Maximum              
License Agreements [Line Items]              
Payments for milestone agreement or earned royalties on achievement of milestones     $ 900        
UCSF License And Option Agreements [Member]              
License Agreements [Line Items]              
Payments for milestone agreement on achievement of milestones           0  
Research and development expense           100 100
Distributed Bio Subscription Agreement [Member]              
License Agreements [Line Items]              
Payments for achievement of specified development and regulatory milestones   $ 5,900          
Research and development expense       $ 100 $ 100 $ 200 $ 100