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Equity Incentive Awards
 9 Months Ended
Sep. 30, 2021
Share-based Payment Arrangement [Abstract]  
Equity Incentive Awards Equity Incentive Awards
Stock Options
The following table summarizes the Company’s option activity and related information:
Number of
Options
(in thousands)		Weighted-
Average
Exercise Price
Balance at December 31, 202010,432 $11.65 
Options granted6,863 8.02 
Options exercised(9)5.86 
Options forfeited(3,427)11.43 
Balance at September 30, 202113,859 $9.91 
Exercisable as of September 30, 20215,724 $10.11 

Restricted Stock Units (“RSUs”)
The following table summarizes the Company’s RSUs activity and related information:
Number of Units
(in thousands)		Weighted-
Average Grant-
Date Fair Value
Outstanding at December 31, 2020540 $6.41 
Granted2,570 3.91 
Vested and released(289)5.35 
Forfeited(228)8.95 
Outstanding at September 30, 20212,593 $3.83 
RSUs granted during the three months ended September 30, 2021 include 1.3 million shares of performance units with both performance and service vesting conditions.
Stock-Based Compensation Expense
The following table presents, by operating expense, the Company’s stock-based compensation expense:
Three Months Ended
September 30,		Nine Months Ended
September 30,
2021202020212020
(In thousands)
Research and development$2,207 $2,051 $7,125 $4,967 
General and administrative3,119 3,969 13,659 9,247 
Total stock-based compensation expense$5,326 $6,020 $20,784 $14,214 
During the nine months ended September 30, 2021, the Company recorded stock-based compensation expenses of approximately $1.6 million, as a result of modifications to certain directors' and officers' stock awards during the periods.